The linear mitochondrial genome of the quarantine chytrid Synchytrium endobioticum : insights into the evolution and recent history of an obligate biotrophic plant pathogen

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creativework.keywords - en
Haplotypes
Soilborne plant pathogens
Potato wart
Plant quarantine
Mitochondrial pathology
Chytridiomycota
Fungal communities
creativework.keywords - fr
Haplotypes
Micro-organismes phytopathogènes du sol
Galle verruqueuse de la pomme de terre
Plantes--Quarantaine
Maladies mitochondriales
Chytridiomycètes
Communautés fongiques
dc.contributor.author
Van de Vossenberg, Bart T. L. H.
Brankovics, Balázs
Nguyen, Hai D. T.
Van Gent-Pelzer, Marga P. E.
Smith, Donna
Dadej, Kasia
Przetakiewicz, Jarosław
Kreuze. Jan F.
Boerma, Margriet
Van Leeuwen, Gerard C. M.
Lévesque, C. André
Van der Lee, Theo A. J.
dc.date.accepted
2018-08-20
dc.date.accessioned
2026-02-03T13:36:39Z
dc.date.available
2026-02-03T13:36:39Z
dc.date.issued
2018-09-10
dc.date.submitted
2018-05-11
dc.description.abstract - en
Background Chytridiomycota species (chytrids) belong to a basal lineage in the fungal kingdom. Inhabiting terrestrial and aquatic environments, most are free-living saprophytes but several species cause important diseases: e.g. Batrachochytrium dendrobatidis, responsible for worldwide amphibian decline; and Synchytrium endobioticum, causing potato wart disease. S. endobioticum has an obligate biotrophic lifestyle and isolates can be further characterized as pathotypes based on their virulence on a differential set of potato cultivars. Quarantine measures have been implemented globally to control the disease and prevent its spread. We used a comparative approach using chytrid mitogenomes to determine taxonomical relationships and to gain insights into the evolution and recent history of introductions of this plant pathogen. Results We assembled and annotated the complete mitochondrial genome of 30 S. endobioticum isolates and generated mitochondrial genomes for five additional chytrid species. The mitochondrial genome of S. endobioticum is linear with terminal inverted repeats which was validated by tailing and PCR amplifying the telomeric ends. Surprisingly, no conservation in organisation and orientation of mitochondrial genes was observed among the Chytridiomycota except for S. endobioticum and its sister species Synchytrium microbalum. However, the mitochondrial genome of S. microbalum is circular and comprises only a third of the 72.9 Kbp found for S. endobioticum suggesting recent linearization and expansion. Four mitochondrial lineages were identified in the S. endobioticum mitochondrial genomes. Several pathotypes occur in different lineages, suggesting that these have emerged independently. In addition, variations for polymorphic sites in the mitochondrial genome of individual isolates were observed demonstrating that S. endobioticum isolates represent a community of different genotypes. Such communities were shown to be complex and stable over time, but we also demonstrate that the use of semi-resistant potato cultivars triggers a rapid shift in the mitochondrial haplotype associated with increased virulence. Conclusions Mitochondrial genomic variation shows that S. endobioticum has been introduced into Europe multiple times, that several pathotypes emerged multiple times, and that isolates represent communities of different genotypes. Our study represents the most comprehensive dataset of chytrid mitogenomes, which provides new insights into the extraordinary dynamics and evolution of mitochondrial genomes involving linearization, expansion and reshuffling. Similar content being viewed by others
dc.identifier.citation
Van de Vossenberg, B. T. L. H., Brankovics, B., Nguyen, H. D. T., Van Gent-Pelzer, M. P. E., Smith, D., Dadej, K., Przetakiewicz, J., Kreuze. J. F., Boerma, M., Van Leeuwen, G. C. M., Lévesque, C. A., & Van der Lee, T. A. J. (2018). The linear mitochondrial genome of the quarantine chytrid Synchytrium endobioticum : insights into the evolution and recent history of an obligate biotrophic plant pathogen. BMC Evolutionary Biology, 18, Article 136. https://doi.org/10.1186/s12862-018-1246-6
dc.identifier.doi
https://doi.org/10.1186/s12862-018-1246-6
dc.identifier.issn
1471-2148
dc.identifier.uri
https://open-science.canada.ca/handle/123456789/4183
dc.language.iso
en
dc.publisher - en
Springer Nature
dc.publisher - fr
Springer Nature
dc.rights - en
Creative Commons Attribution 4.0 International (CC BY 4.0)
dc.rights - fr
Creative Commons Attribution 4.0 International (CC BY 4.0)
dc.rights.uri - en
https://creativecommons.org/licenses/by/4.0/
dc.rights.uri - fr
https://creativecommons.org/licenses/by/4.0/deed.fr
dc.subject - en
Plant diseases
Biology
dc.subject - fr
Maladie des plantes
Biologie
dc.subject.en - en
Plant diseases
Biology
dc.subject.fr - fr
Maladie des plantes
Biologie
dc.title - en
The linear mitochondrial genome of the quarantine chytrid Synchytrium endobioticum : insights into the evolution and recent history of an obligate biotrophic plant pathogen
dc.type - en
Article
dc.type - fr
Article
local.acceptedmanuscript.articlenum
136
local.article.journaltitle - en
BMC Evolutionary Biology
local.article.journalvolume
18
local.pagination
1-15
local.peerreview - en
Yes
local.peerreview - fr
Oui
local.requestdoi - en
No
local.requestdoi - fr
No
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