Additions and corrections: Family-group names in Coleoptera 65 Additions and corrections to “Family-group names in Coleoptera (Insecta)” Patrice Bouchard1, Yves Bousquet2 1 Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, Ontario, K1A 0C6, Canada 2 Gatineau, Quebec, Canada Corresponding author: Patrice Bouchard (Patrice.Bouchard@canada.ca) Academic editor: A. Smith  |  Received 6 September 2019  |  Accepted 15 January 2020  |  Published 25 March 2020 http://zoobank.org/4693D9E0-69DD-4D59-A2E1-B74CA0D26394 Citation: Bouchard P, Bousquet Y (2020) Additions and corrections to “Family-group names in Coleoptera (Insecta)”. ZooKeys 922: 65–139. https://doi.org/10.3897/zookeys.922.46367 Abstract Changes to the treatment of Coleoptera family-group names published by Bouchard et al. (2011) are given. These include necessary additions and corrections based on much-appreciated suggestions from our colleagues, as well as our own research. Our ultimate goal is to assemble a complete list of available Coleoptera family-group names published up to the end of 2010 (including information about their spell- ing, author, year of publication, and type genus). The following 59 available Coleoptera family-group names are based on type genera not included in Bouchard et al. (2011): Prothydrinae Guignot, 1954, Aulonogyrini Ochs, 1953 (Gyrinidae); Po- gonostomini Mandl 1954, Merismoderini Wasmann, 1929, †Escheriidae Kolbe, 1880 (Carabidae); Timarchopsinae Wang, Ponomarenko & Zhang, 2010 (Coptoclavidae); Stictocraniini Jakobson, 1914 (Staphylinidae); Cylindrocaulini Zang, 1905, Kaupiolinae Zang, 1905 (Passalidae); Phaeo- chroinae Kolbe, 1912 (Hybosoridae); Anthypnidae Chalande, 1884 (Glaphyridae); Comophorini Britton, 1957, Comophini Britton, 1978, Chasmidae Streubel, 1846, Mimelidae Theobald, 1882, Rhepsimidae Streubel, 1846, Ometidae Streubel, 1846, Jumnidae Burmeister, 1842, Evambateidae Gistel, 1856 (Scarabaeidae); Protelmidae Jeannel, 1950 (Byrrhoidea); Pseudeucinetini Csiki, 1924 (Limnichidae); Xylotrogidae Schönfeldt, 1887 (Bostrichidae); †Mesernobiinae Engel, 2010, Fab- rasiinae Lawrence & Reichardt, 1966 (Ptinidae); Arhinopini Kirejtshuk & Bouchard, 2018 (Nitid- ulidae); Hypodacninae Dajoz, 1976, Ceuthocera Mannerheim, 1852 (Cerylonidae); Symbiotinae Joy, 1932 (Endomychidae); Cheilomenini Schilder & Schilder, 1928, Veraniini Schilder & Schilder, 1928 (Coccinellidae); Ennearthroninae Chûjô, 1939 (Ciidae); Curtimordini Odnosum, 2010, Mordellochroini Odnosum, 2010 (Mordellidae); Chanopterinae Borchmann, 1915 (Promechei- lidae); Heptaphyllini Prudhomme de Borre, 1886, Olocratarii Baudi di Selve, 1875, Opatrinaires ZooKeys 922: 65–139 (2020) doi: 10.3897/zookeys.922.46367 http://zookeys.pensoft.net Copyright Patrice Bouchard, Yves Bousquet. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. CATALOGUE Launched to accelerate biodiversity research A peer-reviewed open-access journal mailto:Patrice.Bouchard@canada.ca http://zoobank.org/4693D9E0-69DD-4D59-A2E1-B74CA0D26394 https://doi.org/10.3897/zookeys.922.46367 http://zookeys.pensoft.net http://creativecommons.org/licenses/by/4.0/ http://creativecommons.org/licenses/by/4.0/ Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)66 Mulsant & Rey, 1853, Telacianae Poey, 1854, Ancylopominae Pascoe, 1871 (Tenebrionidae); Ox- ycopiini Arnett, 1984 (Oedemeridae); Eutrypteidae Gistel, 1856 (Mycteridae); Pogonocerinae Iablokoff-Khnzorian, 1985 (Pyrochroidae); Amblyderini Desbrochers des Loges, 1899 (Anthicidae); Trotommideini Pic, 1903 (Scraptiidae); Acmaeopsini Della Beffa, 1915, Trigonarthrini Villiers, 1984, Eunidiini Téocchi, Sudre & Jiroux, 2010 (Cerambycidae); Macropleini Lopatin, 1977, Steno- podiides Horn, 1883, Microrhopalides Horn, 1883, Colaphidae Siegel, 1866, Lexiphanini Wilcox, 1954 (Chrysomelidae); †Medmetrioxenoidesini Legalov, 2010, †Megametrioxenoidesini Legalov, 2010 (Nemonychidae); Myrmecinae Tanner, 1966, Tapinotinae Joy, 1932, Acallinae Joy, 1932, Cy- cloderini Hoffmann, 1950, Sthereini Hatch, 1971 (Curculionidae). The following 21 family-group names, listed as unavailable in Bouchard et al. (2011), are determined to be available: Eohomopterinae Wasmann, 1929 (Carabidae); Prosopocoilini Benesh, 1960, Pseu- dodorcini Benesh, 1960, Rhyssonotini Benesh, 1960 (Lucanidae); Galbini Beaulieu, 1919 (Euc- nemidae); Troglopates Mulsant & Rey, 1867 (Melyridae); Hippodamiini Weise, 1885 (Coccinel- lidae); Micrositates Mulsant & Rey, 1854, Héliopathaires Mulsant & Rey, 1854 (Tenebrionidae); Hypasclerini Arnett, 1984; Oxaciini Arnett, 1984 (Oedemeridae); Stilpnonotinae Borchmann, 1936 (Mycteridae); Trogocryptinae Lawrence, 1991 (Salpingidae); Grammopterini Della Beffa, 1915, Aedilinae Perrier, 1893, Anaesthetinae Perrier, 1893 (Cerambycidae); Physonotitae Spaeth, 1942, Octotomides Horn, 1883 (Chrysomelidae); Sympiezopinorum Faust, 1886, Sueinae Muray- ama, 1959, Eccoptopterini Kalshoven, 1959 (Curculionidae). The following names were proposed as new without reference to family-group names based on the same type genus which had been made available at an earlier date: Dineutini Ochs, 1926 (Gyrinidae); Odonteini Shokhin, 2007 (Geotrupidae); Fornaxini Cobos, 1965 (Eucnemidae); Auletobiina Le- galov, 2001 (Attelabidae). The priority of several family-group names, listed as valid in Bouchard et al. (2011), is affected by recent bibliographic discoveries or new nomenclatural interpretations. †Necronectinae Ponomarenko, 1977 is treated as permanently invalid and replaced with †Timarchopsinae Wang, Ponomarenko & Zhang, 2010 (Coptoclavidae); Agathidiini Westwood, 1838 is replaced by the older name Anisoto- mini Horaninow, 1834 (Staphylinidae); Cyrtoscydmini Schaufuss, 1889 is replaced by the older name Stenichnini Fauvel, 1885 (Staphylinidae); Eremazinae Iablokoff-Khnzorian, 1977 is treated as una- vailable and replaced with Eremazinae Stebnicka, 1977 (Scarabaeidae); Coryphocerina Burmeister, 1842 is replaced by the older name Rhomborhinina Westwood, 1842 (Scarabaeidae); Eudysantina Bouchard, Lawrence, Davies & Newton, 2005 is replaced by the older name Dysantina Gebien, 1922 which is not permanently invalid (Tenebrionidae). The names Macraulacinae/-ini Fleutiaux, 1923 (Eucnemidae), Anamorphinae Strohecker, 1953 (Endomychidae), Pachycnemina Laporte, 1840 (Scarabaeidae), Thaumastodinae Champion, 1924 (Limnichidae), Eudicronychinae Girard, 1971 (Elateridae), Trogoxylini Lesne, 1921 (Bostrichidae), Laemophloeidae Ganglbauer, 1899 (Lae- mophloeidae); Ancitini Aurivillius, 1917 (Cerambycidae) and Tropiphorini Marseul, 1863 (Cur- culionidae) are threatened by the discovery of older names; Reversal of Precedence (ICZN 1999: Art. 23.9) or an application to the International Commission on Zoological Nomenclature will be necessary to retain usage of the younger synonyms. Reversal of Precedence is used herein to qualify the following family-group names as nomina protecta: Murmidiinae Jacquelin du Val, 1858 (Cerylonidae) and Cha- lepini Weise, 1910 (Chrysomelidae). The following 17 Coleoptera family-group names (some of which are used as valid) are homonyms of other family-group names in zoology, these cases must be referred to the Commission for a ruling to remove the homonymy: Catiniidae Ponomarenko, 1968 (Catiniidae); Homopterinae Wasmann, 1920, Glyptini Horn, 1881 (Carabidae); Tychini Raffray, 1904, Ocypodina Hatch, 1957 (Staphyli- nidae); Gonatinae Kuwert, 1891 (Passalidae); Aplonychidae Burmeister, 1855 (Scarabaeidae); Mi- Additions and corrections: Family-group names in Coleoptera 67 crochaetini Paulus, 1973 (Byrrhidae); Epiphanini Muona, 1993 (Eucnemidae); Limoniina Jakob- son, 1913 (Elateridae); Ichthyurini Champion, 1915 (Cantharidae); Decamerinae Crowson, 1964 (Trogossitidae); Trichodidae Streubel, 1839 (Cleridae); Monocorynini Miyatake, 1988 (Coc- cinellidae); Gastrophysina Kippenberg, 2010, Chorinini Weise, 1923 (Chrysomelidae); Mecone- mini Pierce, 1930 (Anthribidae). The following new substitute names are proposed: Phoroschizus (to replace Schizophorus Ponomarenko, 1968) and Phoroschizidae (to replace Schizophoridae Ponomarenko, 1968); Mesostyloides (to replace Mesostylus Faust, 1894) and Mesostyloidini (to replace Mesostylini Reitter, 1913). The following new genus-group name synonyms are proposed [valid names in square brackets]: Plo- castes Gistel, 1856 [Aesalus Fabricius, 1801] (Lucanidae); Evambates Gistel, 1856 [Trichius Fabricius, 1775] (Scarabaeidae); Homoeoplastus Gistel, 1856 [Byturus Latreille, 1797] (Byturidae). Two type gen- era previously treated as preoccupied and invalid, Heteroscelis Latreille, 1828 and Dysantes Pascoe, 1869 (Tenebrionidae), are determined to be senior homonyms based on bibliographical research. While Dys- antes is treated as valid here, Reversal of Precedence (ICZN 1999: Art. 23.9) is used to conserve usage of Anomalipus Guérin-Méneville, 1831 over Heteroscelis. Keywords Beetles, family-group name, nomenclature, stem, type genus Introduction Nine years have passed since the publication of the catalogue of “Family-group names in Coleoptera” (Bouchard et al. 2011). During this period, we have become aware of a number of available family-group names that were unfortunately omitted at the time. Furthermore, bibliographic and other errors associated with family-group names and their type genera were discovered. The additions and corrections herein focus on avail- able family-group names proposed up to the end of 2010 with the exception of replace- ment names for names proposed before 2011. Several changes to the classification of Co- leoptera have been published in the years since the publication of Bouchard et al. (2011), generally based on new molecular phylogenetic analyses. A thorough summary of these changes is outside of the scope of this article and therefore the classification scheme used in Bouchard et al. (2011) is followed here to maximize uniformity of contents for us- ers. Further action is required to resolve remaining nomenclatural issues involving the Principle of Priority and the Principle of Homonymy (summarized in Appendices 2, 3). These cases are left unresolved so that specialists on the relevant groups can review them and decide on the best course of action. A dagger symbol “†” precedes fossil taxa. We use the same methods as in Bouchard et al. (2011) although further comments are necessary regarding the treatment of the substantial number of family-group names proposed after 1999 based on the incorrectly formed stem of their type genus. Article 29.4 states that “If after 1999 a new family-group name is based on a generic name which is or ends in a Greek or Latin word or ends in a Greek or Latin suffix, but its derivation does not follow the grammatical procedures of Articles 29.3.1 or 29.3.2, its original spelling must be maintained as the correct original spelling, provided 29.4.1. it Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)68 has a correctly formed suffix [Art. 29.2], and 29.4.2. its stem is formed from the name of the type genus as though it were an arbitrary combination of letters [Art. 29.3.3].” Following the suggestion of Newton (2017: 4), we have accepted that the best way to promote stability in the long term (especially as several more family-group names proposed since 2011 are based on the incorrectly formed stem of their type genus) is to maintain the spellings as originally proposed when conditions laid out in Art. 29.3.3 are met. In this work we have accordingly reverted back to original spellings for family- group names proposed after 1999 that fall into this category. It may be argued that the spelling of some of these family-group names, as used in our initial work (Bouchard et al. 2011), is in prevailing usage and so is to be maintained (Art. 29.5). However, in all the cases the family-group names as corrected in our initial work have been used by very few authors (as far as we known in two works or fewer) and therefore we do not believe that Article 29.5 should apply here. Additions and corrections Bibliographic notes Throughout the text replace “Acloque, 1896” with “Acloque, 1895”. Acloque’s Faune de France was issued in December 1895 (Bousquet 2016: 40), not in 1896 as listed on the title page. Throughout the text replace “Branden, 1885” with “Branden, 1884”. The separate of Branden’s paper in Volume 29 of the Annales de la Société Entomologique de Bel- gique was issued in 1884, prior to the publication in the journal (Bousquet 2016: 543). The paginations in the separate and the journal article are identical. Throughout the text replace “Desmarest, 1857” with “Desmarest, 1852”. We follow Bousquet (2016: 142–143) who recommended using “1852” as the correct date of publication for the second part of this work. Throughout the text replace “Germar, 1824” with “Germar, 1823” (but see exception for the genus Otiorhynchus below). Germar’s Coleopterorum species novae aut minus cognitae” was issued in 1823, not in 1824 as indicated on the title page (Bousquet 2016: 211; Prena 2018: 327, 342). Throughout the text replace “Laporte, 1836” with “Laporte, 1838”. Although 1836 is the year given on the title page of volume 4 of the Revue Entomologique, this vol- ume was only published in early 1838 (Hayek 1983: 207–208). Throughout the text replace “Lacordaire, 1856” with “Lacordaire, 1855”. Lacordaire’s third volume of his Histoire naturelle des insectes was issued in October 1855, not in 1856 as listed on the title page (Bousquet 2016: 314). Throughout the text replace “W. S. MacLeay, 1827” with “W. S. MacLeay, 1826”. MacLeay’s Annulosa. Catalogue of insects, collected by Captain King, R.N. was first published in two volumes in 1826 (Bousquet 2016: 353–354). Additions and corrections: Family-group names in Coleoptera 69 Throughout the text replace “Pic, 1912” with Pic, 1912a.” The family-group names at- tributed to “Pic, 1912” in Bouchard et al. (2011) were published in Coleopterorum Catalogus (18 October 1912) while Ernobiinae (Pic 1912b: 55), which appeared in Volume 28 of L’Échange, was issued earlier in the same year (July 1912). Throughout the text replace “Reitter, 1909” with “Reitter, 1909a.” The family-group names attributed to “Reitter, 1909” in Bouchard et al. (2011) appeared in the second Theil of Reitter’s “Fauna Germanica”, which should be treated as having been published on December 31, 1909 for nomenclatural purposes (recorded in Naturae Novitates in January 1910). The new entry for Tanygnathini Reitter, 1909b (see below) was first proposed in the Coleoptera section (i.e., part 3–4) of “Die Süsswasserfauna Deutschlands: eine Exkursionsfauna” (recorded in Naturae Novitates in August 1909). Table 1 Page 8. In Table 1, replace the generic ending from “-celis” to “-scelis” and the meaning from “spot (Greek)” to “leg (Greek)”. Page 8. In Table 1, replace the meaning of “-onyx” with “claw (Greek)” and that of “-teles” with “end, tail (Greek)”. Catalogue of Coleoptera family-group names Suborder ARCHOSTEMATA Page 95. Below “Suborder ARCHOSTEMATA” add: “Superfamily Cupedoidea Laporte, 1838 Cupesidae Laporte, 1838: 56 [stem: Cuped-]. Type genus: Cupes Fabricius, 1801.” Note. This superfamily is proposed to include the families Crowsoniellidae, Cupedidae, Micromalthidae, Ommatidae, Jurodidae, †Triadocupedidae, †Magnocolei- dae, and †Obrieniidae. The suborder ARCHOSTEMATA includes the superfami- lies Cupedoidea, †Asiocoleoidea, †Rhombocoleoidea, and †Schizocoleoidea. Page 97. Replace the valid name “†Tribe KenderlyKaini Legalov, 2009” with “†Tribe KenderlyKanini Legalov, 2009”. Page 97. In the entry “Kenderlykanini Legalov, 2009c: 285...” replace the stem with “Kenderlykan-” and replace the “Comment” section with “Comment: incorrect original stem formation maintained under Art. 29.4 (should be Kenderlyka-).” Page 97. Move the entries “†Superfamily asioColeoidea Rohdendorf, 1961”, “†Su- perfamily rhomboColeoidea Rohdendorf, 1961” and “†Superfamily sChizo- phoroidea Ponomarenko, 1968” and their associated data to below the entry “Obrieniidae Zherikhin and Gratshev, 1994: 51...” Note. The suborder MYXO- PHAGA includes only the superfamilies Lepiceroidea Hinton, 1936 (1882) and Sphaeriusoidea Erichson, 1845. Page 97. Replace the valid name “†Superfamily sChizophoroidea Ponomarenko, 1968” with “†Superfamily sChizoColeoidea Rohdendorf, 1961”. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)70 Page 97. Replace the entry “Schizophoridae Ponomarenko, 1968: 130…” below the valid name “†Superfamily sChizoColeoidea Rohdendorf, 1961” with: “Schizocoleidae Rohdendorf, 1961: 438 [stem: Schizocole-]. Type genus: Schizoco- leus Rohdendorf, 1961.” Page 97. Replace the valid name “†Family sChizophoridae Ponomarenko, 1968” with “†Family phorosChizidae Bouchard and Bousquet, nomen novum”. Page 97. Replace the entry “Schizophoridae Ponomarenko, 1968: 130…” below the valid name “†Family sChizophoridae Ponomarenko, 1968” with: “Schizophoridae Ponomarenko, 1968: 130 [stem: Schizophor-]. Type genus: Schizo- phorus Ponomarenko, 1968 [preoccupied genus name, not Schizophorus Balashova, 1953 [Trilobita]; syn. of Phoroschizus Bouchard and Bousquet, nomen novum. Comment: permanently invalid (Art. 39): based on preoccupied type genus.” Page 97. Below the entry “Schizophoridae Ponomarenko, 1968: 130…” add: “Phoroschizidae Bouchard and Bousquet, nomen novum for Schizophoridae Ponomarenko, 1968 [stem: Phoroschiz-]. Type genus: Phoroschizus Bouchard and Bousquet, nomen novum for Schizophorus Ponomarenko, 1968.” Page 97. At the end of the entry “Catiniidae Ponomarenko, 1968: 137…” add “Comment: the family-group name Catiniidae Bocquet and Stock, 1957 (type genus Catinia Bocquet and Stock, 1957) is available in Crustacea; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1); the unnecessary replacement name Coleocatiniidae Ponomarenko and Prokin (2015) is unavailable (Art. 11.7.1.1) since it was not based on an available genus name at the time.” Suborder ADEPHAGA Family TriTarsusidae Hong, 2002 Page 99. Replace the valid name “†Family TriTarsidae Hong, 2002” with “†Family TriTarsusidae Hong, 2002”. Page 99. In the entry “Tritarsusidae Hong, 2002: 102...” replace the stem with “Tri- tarsus-” and replace the “Comment” section with “Comment: incorrect original stem formation maintained under Art. 29.4 (should be Tritars-).” Family Gyrinidae Latreille, 1810 Page 100. Replace the valid name “Tribe enhydrini Régimbart, 1882” with “Tribe enhydrusini Branden, 1882”. Page 100. Replace the entry “Enhydrini Régimbart, 1882: 392…” under the valid name “Tribe enhydrusini Branden, 1882” with: “Enhydrini Branden, 1882: 48 [stem: Enhydrus-]. Type genus: Enhydrus Laporte, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1964, 2012a)]. Comment: usage of this name conserved over Dineutini Desmarest, 1851 (Art. 35.5), though considered a junior synonym of Dineutini Desmarest, 1851 by Additions and corrections: Family-group names in Coleoptera 71 Gustafson and Miller (2013: 81, 95–96); name placed on the Official List of Fami- ly-Group Names in Zoology (ICZN 2012a, as “Enhydrusini Régimbart, 1882”).” Page 100. In the entry “Dineutides Desmarest, 1851: 223…” replace “Type genus: Dineutes W. S. MacLeay, 1825.” with “Type genus: Dineutus W. S. MacLeay, 1825.” Page 100. Below the entry “Dineutides Desmarest, 1851: 223…” add: “Dineutini Ochs, 1926: 63 [stem: Dineut-]. Type genus: Dineutus W. S. MacLeay, 1825. Comment: family-group name proposed as new without reference to Di- neutides Desmarest, 1851 (see Gustafson and Miller 2013).” Page 100. Replace the valid name “Subtribe enhydrina Régimbart, 1882” with “Subtribe enhydrusina Branden, 1882”. Page 100. Replace the entry “Enhydrini Régimbart, 1882: 392…” under the valid name “Subtribe enhydrusina Branden, 1882” with: “Enhydrini Branden, 1882: 48 [stem: Enhydrus-]. Type genus: Enhydrus Laporte, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1964, 2012a)]. Comment: correct stem ruled to be Enhydrus- to remove homonymy with Enhydrini Gray, 1825 (type genus Enhydra Fleming, 1822) in Mammalia and “Enhydrusini Régimbart, 1882” placed on the Official List of Family-Group Names in Zoology (ICZN 2012a); “Enhydrini Régimbart, 1882” deemed to be an incorrect original spelling and placed on the Official Index of Rejected and In- valid Family-Group Names in Zoology (ICZN 2012a); regarding the authorship of this name, Régimbart’s paper was issued by December 1882 while Branden’s was published earlier, before 25 April 1882.” Page 100. Below the entry “Enhydrini Branden, 1882: 48…” under the valid name “Subtribe enhydrusina Branden, 1882” add: “Prothydrinae Guignot, 1954: 45 [stem: Prothydr-]. Type genus: Prothydrus Guig- not, 1954.” Page 100. Below the entry “Gyrinites Latreille, 1810: 141…” under the valid name “Subtribe Gyrinina Latreille, 1810” add: “Aulonogyrini Ochs, 1953: 8 [stem: Aulonogyr-]. Type genus: Aulonogyrus Motschul- sky, 1853.” Page 100. Replace the valid name “Tribe oreCToChilini Régimbart, 1882” with “Tribe oreCToChilini Kolbe, 1880”. Page 100. Replace “Orectochilini Régimbart, 1882: 391” with “Orectochilini Kolbe, 1880: 264”. Family Carabidae Latreille, 1802 Page 104. Move the entry “Euryodini W. Horn, 1899: 37…” from “Subtribe dro- miCina Thomson, 1859” to “Subtribe CiCindelina Latreille, 1802” below the entry “Cicindeletae Latreille, 1802: 77…” and replace “Type genus: Euryoda Lacordaire, 1842 [syn. of Prothyma Hope, 1838]” with “Type genus: Euryoda La- cordaire, 1842 [syn. of Heptodonta Hope, 1838]” Note. As discussed by Bousquet (2002: 22) Euryoda Lacordaire, 1842 is an unnecessary replacement name for Hep- todonta Hope, 1838 and is therefore a junior objective synonym of that taxon. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)72 Page 105. Delete the entry “*Colliurides Motschulsky, 1855: 34…” Note. The genus- group name Colliuris used by Latreille (1802) is an incorrect subsequent spelling of Collyris Fabricius, 1801, not in prevailing usage. Therefore, Colliurides used by Motschulsky is an incorrect subsequent spelling of Collyridina Brullé, 1834. Page 105. Below the entry “Ctenostomidae Laporte, 1834b: 38…” add: “Pogonostomini Mandl 1954: 7 [stem: Pogonostomat-]. Type genus: Pogonostoma Klug, 1835. Comment: incorrect original stem formation, not in prevailing usage.” Page 106. In the entry “*Cechenogenici Morawitz, 1889: 40…” replace “Type ge- nus: Cechenus Fischer von Waldheim, 1822” with “Type genus: Cechenus Fischer von Waldheim, 1822 [preoccupied genus name, not Cechenus Illiger, 1807 [Hy- menoptera: Braconidae]; syn. of Cechenochilus Motschulsky, 1850].” Page 106. In the entry “Cechenogenici Csiki, 1927: 110…” replace “Type genus: Cechenus Fischer von Waldheim, 1822” with “Type genus: Cechenus Fischer von Waldheim, 1822 [preoccupied genus name, not Cechenus Illiger, 1807 [Hymenop- tera: Braconidae]; syn. of Cechenochilus Motschulsky, 1850]. Comment: perma- nently invalid (Art. 39): based on preoccupied type genus.” Page 116. In the entry “Sinozolini Deuve, 1997…” replace “Type genus: Sinozolus Bedel, 1898” with “Type genus: Sinozolus Deuve, 1997”. Page 118. Replace the entry “Nomiidae Gozis, 1875: 3…” with: “Nomiidae Gozis, 1875: 3 [stem: Nomius-]. Type genus: Nomius Laporte, 1835 [placed on the Official List of Generic Names in Zoology (ICZN 2011a)]. Com- ment: Nomiidae Gozis, 1875 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 2011a), stem emended to Nomius- and Nomiusidae Gozis, 1875 placed on the Official List of Family-Group Names in Zoology (ICZN 2011a).” Page 119. Below the entry “Carabidomemninae Wasmann, 1928: 271…” add: “Eohomopterinae Wasmann, 1929: 60 [stem: Eohomopter-]. Type genus: Eohomop- terus Wasmann, 1920.” Page 120. At the end of the entry “Homopterinae Wasmann, 1920: 111…” add “Comment: the family-group name Homopterinae Boisduval, 1852 (type genus Homoptera Boisduval, 1852) is available in Lepidoptera; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 120. Below the entry “Paussili Latreille, 1806: 234…” add: “Merismoderini Wasmann, 1929: 60 [stem: Merismoder-]. Type genus: Merismoderus Westwood, 1846 [syn. of Melanospilus Westwood, 1846].” Page 128. At the end of the entry “Glypti G. H. Horn, 1881: 179…” add “Comment: the family-group name Glyptini Cushman and Rohwer, 1920 (type genus Glypta Gravenhorst, 1829) is available and used as valid in Hymenoptera: Ichneumonidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 133. In the entry “Callidides Chaudoir, 1873b: 97…” replace “Type genus: Calleida Dejean, 1824” with “Type genus: Calleida Latreille, 1824”. Page 138. In the entry “Melanodini Alluaud, 1916: 228…” replace “syn. of Mel- anchiton Basilewsky, 1946.” with “syn. of Melanchiton Andrewes, 1940.” Additions and corrections: Family-group names in Coleoptera 73 Page 138. In the entry “Melanchitonitae Jeannel, 1948b: 627…” replace “Type genus: Melanchiton Basilewsky, 1946” with “Type genus: Melanchiton Andrewes, 1940.” Page 141. In the entry “Meleagrosini Morvan, 2004: 2...” replace the stem with “Me- leagros-” and replace the “Comment” section with “Comment: incorrect original stem formation maintained under Art. 29.4 (should be Meleagr-).” Page 143. Replace “Stomidae Chaudoir, 1846: 514” with “Stomides Gené, 1839: 50” Page 145. In the entry “Agronomaeidae Gistel, 1848: [2]…” after “...Amara Bonelli, 1810” add “; preoccupied genus name, not Agronoma Hübner, 1821 [Lepidop- tera]” and at the end of the “Comment” section add “; permanently invalid (Art. 39): based on preoccupied type genus.” Page 146. Above the valid name “Family haliplidae Aubé, 1836” add: “Carabidae incertae sedis †Escheriidae Kolbe, 1880: 266 [stem: Escheri-]. Type genus: Escheria Heer, 1847.” Family CopToClavidae Ponomarenko, 1961 Page 147. Replace the valid name “†Subfamily neCroneCTinae Ponomarenko, 1977” with “†Subfamily TimarChopsinae Wang, Ponomarenko and Zhang, 2010”. Page 147. In the entry “Necronectinae Ponomarenko, 1977: 22…” replace “Type genus: Necronectes Ponomarenko, 1977 [syn. of Timarchopsis Brauer, Redtenbacher and Ganglbauer, 1889]” with “Type genus: Necronectes Ponomarenko, 1977 [pre- occupied genus name, not Necronectes Milne-Edwards, 1881 [Crustacea]; syn. of Timarchopsis Brauer, Redtenbacher and Ganglbauer, 1889]” and add “Comment: permanently invalid (Art. 39): based on preoccupied type genus.” Page 147. Below the entry “Necronectinae Ponomarenko, 1977: 22…” add: “Timarchopsinae Wang, Ponomarenko and Zhang, 2010: 681 [stem: Timarchops-]. Type genus: Timarchopsis Brauer, Redtenbacher and Ganglbauer, 1889. Comment: replacement name for Necronectinae Ponomarenko, 1977; incorrect stem for- mation maintained under Art. 29.4 (should be Timarchopse-).” Page 147. In the entry “Hispanoclavinae Soriano et al., 2007: 527…” replace “Type genus: Hispanoclava Soriano, Ponomarenko and Delclos, 2007” with “Type genus: Hispanoclavina Soriano, Ponomarenko and Delclòs, 2007” and add “Comment: incorrect stem formation maintained under Art. 29.4 (should be Hispanoclavin-).” Family liadyTidae Ponomarenko, 1977 Page 147. In the entry “Liadytidae Ponomarenko, 1977: 37…” replace “Type genus: Liadytes Ponomarenko, 1977” with “Type genus: Liadytes Ponomarenko, 1963.” Family noTeridae Thomson, 1860 Page 148. Replace the valid name “Subfamily noTomiCrinae Zimmermann, 1919” with “Subfamily noTomiCrinae Branden, 1884”. Page 148. Replace “Notomicrini A. Zimmermann, 1919: 110” with “Notomicrini Branden, 1884: 13”. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)74 Family dyTisCidae Leach, 1815 Page 151. In the entry “Bidessini Sharp, 1880: cxlviii…”, replace “Type genus: Bidessus Sharp, 1882” with “Type genus: Bidessus Sharp, 1880 [for comments re- garding problems with the type species of this genus see Bousquet and Bouchard (2018a: 32), Fery and Grygier (2019: 62)].” Page 152. Replace “Hydrocoptini Branden, 1885: 13” with “Hydrocoptini Kolbe, 1883b: 386.” Page 152. Replace “Pachydrini Biström et al., 1997: 66” with “Pachydrini Young, 1980: 306.” Note. We do not consider the statement “should probably be placed in a new tribe” used by Young (1980: 306) as evidence of a conditional proposal (Art. 15.1). Suborder POLYPHAGA Family hydrophilidae Latreille, 1802 Page 157. In the entry “Cyllomina Zaitzev, 1908: 400…” replace “[stem: Cylomat-]” to “[stem: Cylom-]” and delete the “Comment” section. Note. See comment by Seidel et al. (2016: 161) regarding the stem of the genus Cyloma Sharp, 1872. Family hisTeridae Gyllenhal, 1808 Page 160. In the entry “Scolytini Jakobson, 1911a: 652…” replace “Type genus: Sco- lytus Müller, 1764 [preoccupied genus name, not Scolytus Geoffroy, 1762 [Coleop- tera: Curculionidae]; syn. of Onthophilus Leach, 1817]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.” with “Type genus: Scolytus sensu Jakobson, 1911 [not Scolytus Geoffroy, 1762; syn. of Onthophilus Leach, 1817]. Comment: based on a misidentified type genus.” Note. Since stability or universality is not threatened, no application to the Commission is needed to sup- press this family-group name (Art. 65.2.1). Page 161. Replace the valid name “nymphisTrini Tishechkin, 2007” with “nymphis- Terini Tishechkin, 2007”. Page 161. In the entry “Nymphisterini Tishechkin, 2007” replace the stem with “Nymphister-” and replace the “Comment” section with “Comment: incorrect original stem formation maintained under Art. 29.4 (should be Nymphistr-).” Note. See Newton (2017: 5). Family pTiliidae Erichson, 1845 Page 166. In the entry “*Pterycini Dybas, 1966: 16, 44…” replace the “Comment” section with “Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage; the earlier usage of “pterycine group” by Dybas (1955: 562) is unavailable because it is not a noun (Art. 11.7.1.1); also this name has been used subsequently by Hall (2003: 85) but the name Pterycini has not been made available yet (see Hall 2005: 257).” Additions and corrections: Family-group names in Coleoptera 75 Family leiodidae Fleming, 1821 Page 168. Replace the valid name “Tribe aGaThidiini Westwood, 1838” with “Tribe anisoTomini Horaninow, 1834”. Page 168. Below the entry “*Anisotomidae Stephens, 1828: 99…” add: “Anisotomidae Horaninow, 1834: 124 [stem: Anisotom-]. Type genus: Anisotoma Panzer, 1797. Comment: Horaninow (1834) used Anisotoma as valid and in the sense of Panzer (1797).” Page 168. In the entry “Agathidiidae Westwood, 1838: 10…” replace “Type genus: Agathidium Illiger, 1798” with “Type genus: Agathidium Panzer, 1796.” Page 168. Delete the entry “Anisotomidae Reitter, 1884: 6…” Page 169. In the entry “Anisotomidae Erichson, 1845: 41…” delete “; an application will need to be submitted to the Commission to suppress this name for the Prin- ciples of Priority and Homonymy (Art. 65.2.1) if Anisotomidae Reitter, 1884 in Leiodinae: Agathidiini is to be used as valid in the future.” Page 170. Replace the valid names “Tribe anemadini Hatch, 1928” and “Subtribe anemadina Hatch, 1928” with “Tribe anemadini Hatch, 1927” and “Subtribe anemadina Hatch, 1927” respectively. Page 170. Replace “Anemadina Hatch, 1928: 159” with “Anemadina Hatch, 1927: 14” under the valid name “Tribe anemadini Hatch, 1927” and “Subtribe anema- dina Hatch, 1927” respectively. Page 170. At the end of entry “Anemadinae Jeannel, 1936: 179…” replace “Anema- dina Hatch, 1928” with “Anemadina Hatch, 1927”. Page 171. In the entry “Antroherpona Jeannel, 1910: 25...” replace the stem with “Anthroherpon-”. Page 171. In the entry “Antroherpona Guéorguiev, 1974: 841, in key...” replace the stem with “Anthroherpon-” and add at the end of the “Comment” section “incor- rect original stem formation, not in prevailing usage.” Family sTaphylinidae Latreille, 1802 Page 175. In the entry “Omalidae W. S. MacLeay, 1825…” replace “Type genus: Omalium Gravenhorst, 1802” with “Type genus: Omalium Gravenhorst, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2015b)].” Page 176. In the entry “Omalidae W. S. MacLeay, 1825…” replace “Type genus: Omalium Gravenhorst, 1802” with “Type genus: Omalium Gravenhorst, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2015b)].” and at the end of the “Comment” section add: “; name placed on the Official List of Family-Group Names in Zoology (ICZN 2015b; as Omaliidae MacLeay, 1825)”. Page 178. In the entry “Megarthrini Joy, 1932: 93…” replace “Type genus: Megar- thrus Curtis, 1829” with “Type genus: Megarthrus Stephens, 1829”. Page 181. In the entry “Neocerini Jeannel, 1954a: 316…” add the following at the end of the “Comment” section: “; the older family-group name Neocerini Salmon, 1941 (type genus Neocerus Salmon, 1941, preoccupied genus name, not Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)76 Neocerus Wasmann, 1893) is available though permanently invalid (Art. 39) in Collembola.” Page 182. Replace the valid name “Tribe Jubini Raffray, 1904” with “Tribe Jubini Raffray, 1898”. Page 182. Replace “Jubini Raffray, 1904: 489, in key” with “Jubini Raffray, 1898: 199”. Page 183. Replace the valid name “Tribe mayeTiini Winkler, 1925” with “Tribe may- eTiini Scheerpeltz, 1925”. Page 183. Replace “Mayetiini Winkler, 1925: 348” with “Mayetiini Scheerpeltz, 1925: 348.” Note. The section on Staphylinidae, except for the Steninae (pp. 349–357), Euaesthetinae (pp. 357–358) and the genus Staphylinus (pp. 381–386), was compiled (“Conscripsit”) by Scheerpeltz (p. 323) and therefore he should be treated as the author of the new taxa in this section. Page 184. Replace the valid name “Subtribe Trimiina Bowman, 1934” with “Sub- tribe Trimiina Brendel and Wickham, 1890”. Page 184. Replace “Trimii Bowman, 1934: 8” with “Trimiini Brendel and Wickham, 1890: 225”. Page 184. In the entry “Trimiina Jeannel, 1950a: 139…” replace “without reference to Trimii Bowman, 1934” with “without reference to Trimii Brendel and Wickham, 1890”. Page 184. Replace the valid names “Tribe TroGasTrini Jeannel, 1949” and “Subtribe TroGasTrina Jeannel, 1949” with “Tribe TroGasTrini Brendel and Wickham, 1890” and “Subtribe TroGasTrina Brendel and Wickham, 1890” respectively. Page 184. Below the valid names “Tribe TroGasTrini Jeannel, 1949” and “Subtribe TroGasTrina Jeannel, 1949” replace “Trogastrini Jeannel, 1949a: 41, in key” with “Trogasterini Brendel and Wickham, 1890: 225” and add “Comment: in- correct stem formation, not in prevailing usage.” Page 186. In the entry “Bythinini Raffray, 1890: 83, in key…” under the valid names “Tribe byThinini Raffray, 1890” and “Subtribe byThinina Raffray, 1890” add “Comment: published March 1890; this family-group name was also proposed in the same year by Brendel and Wickham (1890 [June]: 224, as Bythinini).” Page 188. At the end of the entry “Tychini Raffray, 1904: 490, in key…” add “Com- ment: junior homonym of Tychinae Dana, 1851 (type genus: Tyche Bell, 1835), which has been used as valid in Crustacea: Decapoda recently (e.g., Davie et al. 2015); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 193. In the entry “Athetae Casey, 1910: 2…” under the valid names “Tribe aTheTini Casey, 1910” and “Subtribe aTheTina Casey, 1910” replace the “Comment” section with “Comment: name placed on the Official List of Fam- ily-Group Names in Zoology (ICZN 2012e).” Note. The name “Athetae” used earlier by Rambousek (1907: 40) is considered a plural term for subgenera of Atheta, Rambousek’s name is therefore unavailable as a family-group name (Art. 11.7.1.2). Additions and corrections: Family-group names in Coleoptera 77 Page 193. In the entry “Callicerina Jakobson, 1908: 448…” replace “Type genus: Cal- licerus Gravenhorst, 1802” with “Type genus: Callicerus Gravenhorst, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2012e)].” and replace the “Comment” section with “Comment: following an application by Gusarov (2011) the names Callicerini Jakobson, 1908, Callicerini Horion, 1967 and Callicerini Lohse, 1969, which are junior homonyms of Callicerini Rondani, 1856 (type genus Callicera Panzer, 1806) in Diptera: Syrphidae, were suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy and placed on the Of- ficial Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 2012e).” Page 194. At the end of the entry “Geostibae Seevers, 1978: 126…” add “Comment: following an application by Gusarov (2011) the name Geostibina Seevers, 1978 was placed on the Official List of Family-Group Names in Zoology (ICZN 2012e).” Page 195. In the entries “Corotocini Fenyes, 1918: 61…” replace “Type genus: Coro- toca Schiødte, 1847” with “Type genus: Corotoca Schiødte, 1853”. Page 197. Replace the valid name “CrypTonoTopseini Pace, 2003” with “CrypTono- Topsisini Pace, 2003”. Page 197. In the entry “Cryptonotopsisini Pace, 2003: 38...” replace the stem with “Cryptonotopsis-” and replace the “Comment” section with “Comment: incorrect original stem formation maintained under Art. 29.4 (should be Cryptonotopse-).” Note. See Newton (2017: 5). Page 203. Replace the valid names “Tribe mimeCiTini Wasmann, 1917” and “Sub- tribe mimeCiTina Wasmann, 1917” with “Tribe mimeCiTini Wasmann, 1909” and “Subtribe mimeCiTina Wasmann, 1909” respectively. Page 203. Replace “Mimecitonini Wasmann, 1917: 325” with “Mimecitonini Was- mann, 1909: 55”. Page 205. Replace the valid name “Tribe oxypodinini Fenyes, 1921” with “Tribe oxypodinini Fenyes, 1918”. Page 210. In the entry “Prognathites Blanchard, 1845a: 290…” replace “Type ge- nus: Prognathus Berthold, 1827” with “Type genus: Prognathus Blondel, 1827”. Page 215. In the entry “Chevrolatini Reitter, 1882c: 142…” replace “Type genus: Chev- rolatia Jacquelin du Val, 1859” with “Type genus: Chevrolatia Jacquelin du Val, 1850”. Page 215. Replace the valid name “Tribe CyrTosCydmini Schaufuss, 1889” with “Tribe sTeniChnini Fauvel, 1885” and move names for the entire tribe down to just above the valid name “Subfamily sTeninae MacLeay, 1825” to maintain al- phabetical order of valid tribes. Note. The name Cyrtoscydmini Schaufuss, 1889 was replaced by Glandulariini Schaufuss, 1889 by Newton (2015) but Glan- dulariini needs to be replaced by Stenichnini Fauvel, 1885 based on priority. Page 215. Replace “Stenichnini Ganglbauer, 1898: 25” with “Stenichnini Fauvel, 1885: 182” and move the entry “Stenichnini Fauvel, 1885: 182…” above to im- mediately under the valid name “Tribe sTeniChnini Fauvel, 1885”. Page 216. In the entry “Austroaesthetini Cameron, 1944: 69...” replace “as Austroaes- thetus, unjustifed emendation of genus name not in prevailing usage” with “as Aus- troaesthetus, incorrect subsequent spelling of genus name not in prevailing usage”. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)78 Page 216. Replace the valid name “Tribe Fenderiini Scheerpeltz, 1974” with “Tribe sTiCToCraniini Jakobson, 1914”. Page 216. Above the entry “Fenderiini Scheerpeltz, 1974: 103...” add: “Stictocraniini Jakobson, 1914: 529 [stem: Stictocrani-]. Type genus: Stictocranius J. L. LeConte, 1866.” Note. See Newton (2017: 5). Page 223. In the entry “Philonthidae Kirby, 1837: 91…” replace “Type genus: Philonthus Curtis, 1829” with “Type genus: Philonthus Stephens, 1829”. Page 224. In the entry “Ocypina Hatch, 1957: 173, in key…” replace the “Comment” section with “Comment: incorrect original stem formation, not in prevailing us- age; usage of the corrected stem Ocypod- proposed by Newton and Thayer (1992: 65) creates a homonymy problem with Ocypodidae Rafinesque, 1815 (type ge- nus Ocypode Weber, 1795), which has been used as valid in Crustacea (e.g., Nad- erloo 2017), and therefore this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 224. Replace “Tanygnathinini Reitter, 1909: 105” with “Tanygnathinini Reit- ter, 1909b: 164”. Series SCARABAEIFORMIA Page 225. In the entry “Scarabaeïdes Latreille, 1802…” replace “Type genus: Scara- baeus Linnaeus, 1758.” with “Type genus: Scarabaeus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 2014)].” Family GeoTrupidae Latreille, 1802 Page 226. Replace the valid name “Tribe bolbelasmini Nikolajev, 1996” with “Tribe bolbelasmini Iablokoff-Khnzorian, 1977”. Page 226. Replace “Bolbelasmini Nikolajev, 1996: 96” with “Bolbelasmini Iablokoff-Khnzorian, 1977: 165”. Page 227. Replace the valid name “Tribe odonTeini Shokhin, 2007” with “Tribe odonTeini Streubel, 1846”. Page 227. Above the entry “Odonteini Shokhin, 2007: 111…” add: “Odontaeidae Streubel, 1846: 960 [stem: Odonte-]. Type genus: Odonteus Samouelle, 1819 [as Odontaeus, incorrect subsequent spelling of type genus name, not in pre- vailing usage; placed on the Official List of Generic Names in Zoology (ICZN 2006a)]. Comment: incorrect original stem formation, not in prevailing usage.” Page 227. In the entry “Odonteini Shokhin, 2007: 111…” replace “…Bolboceras Kirby, 1819 was fixed differently by the Commission (ICZN 2006a).” with “…Bolboceras Kirby, 1819 was fixed differently by the Commission (ICZN 2006a); family-group name proposed as new without reference to Odonteini Streubel, 1846.” Family passalidae Leach, 1815 Page 228. In the entry “Ceracupini Boucher, 2006: 319...” replace the stem with “Ceracup-” and replace the “Comment” section with “Comment: incorrect origi- nal stem formation maintained under Art. 29.4 (should be Ceracuped-).” Additions and corrections: Family-group names in Coleoptera 79 Page 228. Below the entry “Ceracupedini Boucher, 1006: 319...” add: “Tribe CylindroCaulini Zang, 1905 Cylindrocaulinae Zang, 1905: 229 [stem: Cylindrocaul-]. Type genus: Cylindrocau- lus Fairmaire, 1880.” Note. Boucher et al. (2017) recently proposed the family- group name Ceracyclini for the genera Cylindrocaulus Fairmaire, 1880 and the new fossil genus Ceracyclus. However, Cylindrocaulini Zang, 1905 is older and should be used as valid instead of Ceracyclini. Page 229. At the end of the entry “Gonatinae Kuwert, 1891: 169…” add “Com- ment: the family-group name Gonatidae Hoyle, 1886 (type genus: Gonatus Gray, 1849) is available in Cephalopoda; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 229. In the entry “Pelopinae Kuwert, 1896: 229…” replace “… [stem: Pelopid-]. Type genus: Pelopides Kuwert, 1896. Comment: incorrect original stem forma- tion, not in prevailing usage.” with “…[stem: Pelop-]. Type genus: Pelops Kaup, 1871 [preoccupied genus name, not Pelops Koch, 1835 [Acari: Oribatida], syn. of Protomocoelus Zhang, 1905]. Comment: the older genus group-name Pelops Gistel, 1834 [Coleoptera: Tenebrionidae] was recently treated as a nomen oblitum by Bousquet and Bouchard (2017: 132); permanently invalid (Art. 39), based on preoccupied type genus.” Page 229. Below the entry “Pleurariinae Kuwert, 1896: 224...” add: “Kaupiolinae Zang, 1905: 227 [stem: Kaupiol-]. Type genus: Kaupiolus Zang, 1903 [syn. of Labienus Kaup, 1871].” Page 229. Replace “Gnaphalocneminae Gravely, 1914: 194” with “Gnaphalocne- minae Heller, 1900: 11”. Family Glaresidae Prudhomme de Borre, 1886 Page 231. Replace the valid name “Family Glaresidae Kolbe, 1905” with “Family Glaresidae Prudhomme de Borre, 1886”. Page 231. Replace the entry “Glaresini Kolbe, 1905: 543...” with: “Glaresini Prud- homme de Borre, 1886: 56 [stem: Glares-]. Type genus: Glaresis Erichson, 1848.” Family luCanidae Latreille, 1804 Page 232. Below the entry “Aesalidae W. S. MacLeay, 1819: 102…” under the valid name “Tribe aesalini MacLeay, 1819” add: “Plocasteidae Gistel, 1856a: 365 [stem = Plocast-]. Type genus: Plocastes Gistel, 1856 [Gistel (1856a: 365) included one species under the generic name Plocastes, scar- aboides, which refers to Lucanus scarabaeoides Panzer, 1795 and this species is the type species by monotypy; syn. nov. of Aesalus Fabricius, 1801]. Comment: incor- rect original stem formation, not in prevailing usage.” Page 234. Replace the entry “*Chalcodinae Didier and Séguy, 1953: 91…” with: “Chalcodinae Didier and Séguy, 1953: 12, in key, 91 [stem: Chalcod-]. Type species: Chalcodes H. C. C. Burmeister, 1847.” Note. This family-group name is available since the authors provided a description in the key. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)80 Page 234. In the entry “*Prosopocoilini Benesh, 1960: 50...” remove the asterisk (*) and replace the “Comment” section with “Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Klausnitzer (1995: 10, as Prosopocoilini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (Art. 13.2.1).” Page 234. In the entry “*Pseudodorcini Benesh, 1960: 97...” remove the asterisk (*) and replace the “Comment” section with “Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Klausnitzer (1995: 10, as Pseudodorcini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (Art. 13.2.1).” Page 234. In the entry “*Rhyssonotini Benesh, 1960: 148...” remove the asterisk (*) and replace the “Comment” section with “Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Klausnitzer (1995: 10, as Rhyssonotini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (Art. 13.2.1); incorrect original stem formation, not in prevailing usage.” Page 234. Delete the entry “*Chalcodinae J. P. Lacroix, 1979: 258…” Family oChodaeidae Mulsant and Rey, 1871 Page 235. Replace the valid names “Family oChodaeidae Mulsant and Rey, 1871”, “Subfamily oChodaeinae Mulsant and Rey, 1871” and “Tribe oChodaeini Mulsant and Rey, 1871” with “Family oChodaeidae Streubel, 1846”, “Sub- family oChodaeinae Streubel, 1846” and “Tribe oChodaeini Streubel, 1846” respectively. Page 235. Below the valid names “Family oChodaeidae Streubel, 1846”, “Subfamily oChodaeinae Streubel, 1846” and “Tribe oChodaeini Streubel, 1846” replace the entry “Ochodéens Mulsant and Rey, 1871b: 493…” with: “Ochodaeidae Streubel, 1846: 960 [stem: Ochodae-]. Type genus: Ochodaeus Dejean, 1821.” Family hybosoridae Erichson, 1847 Page 237. Replace the entry “Acanthocérides Lacordaire, 1856: 155…” with: “Acanthoceridae Streubel, 1846: 960 [stem: Acanthocer-]. Type genus: Acanthocerus W. S. MacLeay, 1819 [preoccupied genus name, not Acanthocerus Palisot de Beau- vois, 1818 [Hemiptera]; syn. of Ceratocanthus A. White, 1842]. Comment: per- manently invalid (Art. 39): based on preoccupied type genus.” Page 237. In the entry “Ceratocanthini Martínez, 1968: 14…” replace “Acantho- cerini Lacordaire, 1856” with “Acanthocerini Streubel, 1846”. Page 237. Below the entry “Hybosoridae Erichson, 1847a: 104...” add: “Phaeo- chroinen Kolbe, 1912: 153 [stem: Phaeochro-]. Type genus: Phaeochrous Laporte, Additions and corrections: Family-group names in Coleoptera 81 1840. Comment: name listed as “Phaeochroinae” in the Sachregister (p. v), like- ly written by the editor Karl Grünberg.” Family Glaphyridae MacLeay, 1819 Page 238. Below the entry “Glaphyridae W. S. MacLeay, 1819: 76…” add: “Anthypnidae Chalande, 1884: 46, 99 [stem= Anthypn-]. Type genus: Anthypna Eschscholtz, 1818.” Family sCarabaeidae Latreille, 1802 Page 238. In the entry “Scarabaeïdes Latreille, 1802…” replace “Type genus: Scara- baeus Linnaeus, 1758.” with “Type genus: Scarabaeus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 2014)].” Page 239. Replace the valid name “Subfamily eremazinae Iablokoff-Khnzorian, 1977” with “Subfamily eremazinae Stebnicka, 1977”. Page 239. Replace the entry “Eremazini Iablokoff-Khnzorian, 1977 [3 October]: 168…” with: “*Eremazini Iablokoff-Khnzorian, 1977 [3 October]: 168 [stem: Eremaz-]. Type ge- nus: Eremazus Mulsant, 1851. Comment: unavailable family-group name, pro- posed after 1930 without description or bibliographic reference to such a descrip- tion (Art. 13.1).” Note. The only descriptive terms provided for this name are “Bei den ErEmizi ist das Metendosternit wie bei den AphodiinAE gebildet, denen sie viel näher stehen als die anderen AEgiAlitinAE, auch ihrer Gesamtbildung nach” [In the case of the Eremizi the metendosternite is formed as in the Aphodiinae, to which they stand much closer than the other Aegialitinae, also in their general struc- ture]. This statement does not fulfill article 13.1.1, which states “be accompanied by a description or definition that states in words characters that are purported to differentiate the taxon.” Therefore this name is unavailable. Page 239. In the entry “Eremazina Stebnicka, 1977 [“31 December”]: 412…” delete the “Comment” section. Page 239. In the entry “Aphodida Leach, 1815: 97…” below the valid names “Sub- family aphodiinae Leach, 1815”, “Tribe aphodiini Leach, 1815” and “Subtribe aphodiina Leach, 1815” replace “Type genus: Aphodius Illiger, 1798” with “Type genus: Aphodius Hellwig, 1798.” Note. See Alonso-Zarazaga and Krell (2011). Page 242. In the entry “Scarabaeïdes Latreille, 1802…” replace “Type genus: Scara- baeus Linnaeus, 1758.” with “Type genus: Scarabaeus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 2014)].” Page 243. Replace the valid name “Tribe Gymnopleurini Lacordaire, 1856” with “Tribe Gymnopleurini Streubel, 1846”. Page 243. Replace the entry “Gymnopleurides Lacordaire, 1856: 72…” with: “Gymnopleuridae Streubel, 1846: 961 [stem: Gymnopleur-]. Type genus: Gymnopleu- rus Illiger, 1803.” Page 244. Replace the valid name “Tribe onThophaGini Burmeister, 1846” with “Tribe onThophaGini Streubel, 1846”. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)82 Page 244. Replace the entry “Onthophagidae H. C. C. Burmeister, 1846: [1]…” with: “Onthophagidae Streubel, 1846: 961 [stem: Onthophag-]. Type genus: Onthophagus Latreille, 1802. Comment: published by 13 August 1846; this family-group name was also used in the same year by Burmeister (1846 [by 26 November 1846]: [1], as Onthophagidae).” Page 245. In the entry “Scarabaeïdes Latreille, 1802…” replace “Type genus: Scara- baeus Linnaeus, 1758.” with “Type genus: Scarabaeus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 2014)].” and at the end add: “Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 2014, as Scarabaeidae Latreille, 1802).” Page 248. Replace the valid name “Tribe Comophorinini Britton, 1957” with “Tribe Comophini Britton, 1978”. Page 248. Replace the entry “Comophorini Britton, 1957: 10…” with the following entries: “Comophorini Britton, 1957: 10 [stem: Comophor-]. Type genus: Comophorus Blan- chard, 1850 [preoccupied genus name, not Comophorus Agassiz, 1846 [Pisces]; syn. of Comophorina Strand, 1928]. Comment: permanently invalid (Art. 39): based on preoccupied type genus. Comophini Britton, 1978: 7 [stem: Comoph-]. Type genus: Comophus Britton, 1978 [syn. of Comophorina Strand, 1928]. Comment: replacement name for Como- phorini Britton, 1957 because of the homonymy of the type genus. Comophorinini Britton, 1987: 761 [stem: Comophorin-]. Type genus: Comophorina Strand, 1928.” Page 249. In the entry “Hétéronycides Lacordaire, 1856: 225...” at the end of the “Comment” section add “; the family-group name Heteronychidae André, 1891 (type genus Heteronyx Saussure, 1887) is available in Hymenoptera though perma- nently invalid (based on preoccupied type genus).” Page 250. Below the entry “Lepisiidae H. C. C. Burmeister, 1844: 166…” add: “Chasmidae Streubel, 1846: 960 [stem: Chasm-]. Type genus: Chasme Lepeletier and Audinet-Serville, 1828.” Page 250. Replace the entry “Lepitrichiden Oken, 1843: 483…” with: “Lepitrichina Perty, 1840: 933 [stem: Lepitrich-]. Type genus: Lepitrix Lepeletier and Audinet-Serville, 1828.” Note. The family-group name Lepitrichina Perty, 1840, published by 17 November 1840 (Bousquet 2016: 413), is a senior synonym of Pachycnemina Laporte, 1840, issued by 26 December 1840 (Bousquet 2016: 321). Reversal of Precedence (ICZN 1999: Article 23.9) or an application to the Commission is necessary to conserve usage of Pachycnemina Laporte as valid. Page 250. Replace the entry “Haplonychidae H. C. C. Burmeister, 1855: 224…” with: “Haplonychidae H. C. C. Burmeister, 1855: 224 [stem: Aplonych-]. Type genus: Aplonycha Boisduval, 1835 [as Haplonycha, unjustifed emendation of type genus name by Agassiz (1846b: 29)]. Comment: the unjustified emendation Haplo- nycha Agassiz, 1846 is in prevailing usage but attributed to Dejean (1836), not to Boisduval (1835) who first made the name Aplonycha available and therefore, Additions and corrections: Family-group names in Coleoptera 83 Art. 33.2.3.1 (ICZN 1999) cannot be used to consider Haplonycha as a justified emendation; incorrect original stem formation, not in prevailing usage; the junior homonym Aplonychini De Stefani, 1908 (type genus Aplonyx De Stefani, 1908) is available in Diptera; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 252. In the entry “Rhizotrogidae H. C. C. Burmeister, 1855: 308…”, replace “Type genus: Rhizotrogus Latreille, 1825” with “Type genus: Rhizotrogus Berthold, 1827” Page 257. Below the entry “Phyllurgaeidae Gistel, 1848: [5]…” add: “Mimelidae Theobald, 1882: 112 [stem = Mimel-]. Type genus: Mimela Kirby, 1823.” Page 257. Replace the valid name “Subtribe popilliina Ohaus, 1918” with “Subtribe popilliina Ohaus, 1902”. Page 257. Replace “Popilliina Ohaus, 1918: 133” with “Popilliidae Ohaus, 1902: 270” Page 258. Below the entry “Anoplognathidae W. S. MacLeay, 1819: 81…” under the valid name “Subtribe anoploGnaThina MacLeay, 1819” add: “Rhepsimidae Streubel, 1846: 960 [stem: Repsim-]. Type genus: Repsimus W. S. Mac- Leay, 1819. Comment: incorrect original stem formation, not in prevailing usage.” Page 259. Replace “Macraspididae H. C. C. Burmeister, 1844: 343” with “Mac- raspida Perty, 1840: 933”. Page 259. Below the entry “Pelidnotidae H. C. C. Burmeister, 1844: 388…” add: “Ometidae Streubel, 1846: 960 [stem: Omet-]. Type genus: Ometis Latreille, 1829 [syn. of Lagochile Hoffmannsegg, 1817].” Page 260. In the entry “Dynastidae W. S. MacLeay, 1819…” under the valid name “Subfamily dynasTinae MacLeay, 1819” replace “Type genus: Dynastes W. S. MacLeay, 1819.” with “Type genus: Dynastes W. S. MacLeay, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 2014)].” and in the same entry under the valid name “Tribe dynasTini MacLeay, 1819” replace “Type genus: Dynastes W. S. MacLeay, 1819.” with “Type genus: Dynastes W. S. MacLeay, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 2014)].” and at the end add: “Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 2014).” Page 260. Replace “Chalepidae H. C. C. Burmeister, 1847: 71” with “Chalepidae Streubel, 1846: 960”. Page 261. In the entry “Oryctésaires Mulsant, 1842: 372…” replace “Type genus: Oryctes Illiger, 1798” with “Type genus: Oryctes Hellwig, 1798.” Note. See Alonso- Zarazaga and Krell (2011). Page 264. In the entry “Aspilina Krikken, 1984: 25, in key…” replace “Type genus: Aspilus Westwood in Schaum, 1848” with “Type genus: Aspilus Schaum, 1848” Page 265. In the entry “Spilophorina Krikken, 1984: 25, in key…” replace “Type genus: Spilophorus Westwood in Schaum, 1848” with “Type genus: Spilophorus Schaum, 1848”. Page 265. In the entry “Trogodina Krikken, 1984: 27, in key…” replace “Type ge- nus: Trogodes Westwood, 1874” with “Type genus: Trogodes Boheman, 1851”. Page 266. Replace the valid name “Subtribe CoryphoCerina Burmeister, 1842” with “Subtribe rhomborhinina Westwood, 1842”. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)84 Page 266. In the entry “Coryphoceridae H. C. C. Burmeister, 1842: 215...” add “Comment: published by 8 December 1842.” Page 266. Below the entry “Coryphoceridae H. C. C. Burmeister, 1842: 215...” add: “Jumnidae H. C. C. Burmeister, 1842: 195 [stem: Jumn-]. Type genus: Jumnos Saun- ders, 1839. Comment: published by 8 December 1842; we act as First Revisers (Coryphoceridae H. C. C. Burmeister, 1842 vs Jumnidae H. C. C. Burmeister, 1842) and select Coryphoceridae H. C. C. Burmeister, 1842 to have precedence. Page 266. Replace the entry “Rhomborrhinae Shoch, 1894: 170…” with: “Rhomborrhinae Westwood, 1842: 340 [stem: Rhomborhin-]. Type genus: Rhom- borhina Hope, 1837 [as Rhomborrhina, incorrect subsequent spelling of the type genus, not in prevailing usage]. Comment: published on 1 January 1842; incorrect original stem formation, not in prevailing usage.” and move entire entry above “Coryphoceridae H. C. C. Burmeister, 1842: 215...” to keep the chronological order. Note. We have decided to use Rhomborhinina Westwood, 1842 as valid (as determined by the Principle of Priority) based on the fact that Cetoniinae taxonomy is in flux and many changes to the classification will be necessary in the future (A. B. T. Smith, pers. comm. September 2019). Page 271. Below the entry “Paniscidae Gistel, 1848: [5]…” add: “Evambateidae Gistel, 1856a: 365 [stem = Evambat-]. Type genus: Evambates Gistel, 1856 [Gistel (1856a: 365) originally included Scarabaeus fasciatus Linnaeus, 1758 and Trichius zonatus Germar, 1831 in his genus Evambates; we hereby select Scara- baeus fasciatus Linnaeus, 1758 as the type species of Evambates Gistel, 1856; syn. nov. of Trichius Fabricius, 1775]. Comment: incorrect original stem formation, not in prevailing usage.” Page 271. In the entry “Myodermini Péringuey, 1907: 313…” replace “[stem: Myoderm-]. Type genus: Myodermum H. C. C. Burmeister and Schaum, 1840.” with “[stem: My- odermat-]. Type genus: Myoderma Dejean, 1833. Comment: incorrect original stem formation, not in prevailing usage.” Note. See Bousquet and Bouchard (2013: 38) for comments regarding the correct spelling and authorship of the type genus. Family euCineTidae Lacordaire, 1857 Page 273. In the entry “Cryptomeridae Broun, 1893: 1358…” replace “Type genus: Cryptomera Broun, 1893 [syn. of Eucinetus Germar, 1818]” with “Type genus: Cryptomera Broun, 1893 [preoccupied genus name, not Cryptomera Rafinesque, 1820 [Chilopoda]; syn. of Eucinetus Germar, 1818]. Comment: permanently in- valid (Art. 39): based on preoccupied type genus.” Family bupresTidae Leach, 1815 Page 278. In the entry “Xenopsina Volkovitsh, 2008: 628...” replace the stem with “Xenops-” and replace the “Comment” section with “Comment: incorrect original stem formation maintained under Art. 29.4 (should be Xenopse-).” Page 281. In the entry “Polybothrisidae Gistel, 1848…” replace “Type genus: Poly- bothris Spinola, 1837.” with “Type genus: Polybothris Dupont, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 2015a)].” Additions and corrections: Family-group names in Coleoptera 85 Page 281. Replace “Capnodina Jakobson, 1913: 779” with “Capnodini Csiki, 1909c: 168”. Page 285. In the entry “Melobasini Bílý, 2000: 113...” replace the stem with “Melobas-” and replace the “Comment” section with “Comment: incorrect original stem for- mation maintained under Art. 29.4 (should be Melobase-).” Page 288. Under the valid name “Subtribe aphanisTiCina Jacquelin du Val, 1859” in the entry “Aphanisticites Jacquelin du Val, 1859: 104…” replace “Type genus: Aphanisticus Latreille, 1829” with “Type genus: Aphanisticus Latreille, 1810.” Note. The genus Aphanisticus is often credited in the literature to La- treille (1829a: 448). However, the name was first made available by Latreille (1810: 169). Page 288. Replace the valid name “Subtribe amorphosomaTina Majer, 2000” with “Subtribe amorphosomina Majer, 2000”. Page 288. In the entry “Amorphosomina Majer, 2000: 210...” replace the stem with “Amorphosom-” and replace the “Comment” section with “Comment: incorrect original stem formation maintained under Art. 29.4 (should be Amorphosomat-).” Page 289. Replace the valid name “Subtribe ClemaTina Majer, 2000” with “Subtribe Clemina Majer, 2000”. Page 289. In the entry “Clemina Majer, 2000: 215...” replace the stem with “Clem-” and replace the “Comment” section with “Comment: incorrect original stem for- mation maintained under Art. 29.4 (should be Clemat-).” Family byrrhidae Latreille, 1804 Page 291. Replace “Lioonini Leng, 1920: 193” with “Lioonini Casey, 1912: 59”. Page 291. At the end of the entry “Microchaetini Paulus, 1973: 353, in key…” add “Comment: the senior homonym Microchaetini Beddard, 1895 (type genus Mi- crochaetus Rapp, 1849) in Oligochaeta is currently used as valid (see Plisko 2013: 85); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Superfamily byrrhoidea Latreille, 1804 Page 292. Above the valid name “Family elmidae Curtis, 1830” add: “Family proTelmidae Jeannel, 1950 Protelmini Jeannel, 1950: 170, in key [stem: Protelm-]. Type genus: Protelmis Grou- velle, 1911.” Family elmidae Curtis, 1830 Page 293. Replace the entry “Limniidae Hope 1838a: 153…” with: “Limniidae C. G. Thomson, 1859: 21 [stem: Limni-]. Type genus: Limnius Illiger, 1802.” Note. Limniidae Hope, 1838a: 153 is unavailable since it was not based on a genus used as valid at the time. Hope (1838a) listed Limnius (as “Limneus”) as a synonym of Stenelmis Dufour, 1835. Family limniChidae Erichson, 1846 Page 295. Above the entry “Thaumastodinae Champion, 1924: 25…” add: Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)86 “Pseudeucinetini Csiki, 1924: 14 [stem: Pseudeucinet-]. Type genus: Pseudeucinetus Heller, 1921. Comment: the names Thaumastodinae Champion and Pseudeuci- netini Csiki were published in the same year, Csiki’s publication was issued on 24 February 1924 (verso of title page) while that of Champion was published in the Feb- ruary 1924 issue of the journal The Entomologist’s Monthly Magazine; Article 35.5 of the Code (ICZN 1999), proposed by Hernando and Ribera (2016: 32) as a reason to preserve usage of the subfamily name Thaumastodinae, cannot be used in this case, the main reason is that Thaumastodini and Pseudeucinetini are not, and cannot, be used as two separate valid tribes within the subfamily Thaumastodinae since they are based on a single taxonomic entity (i.e., the genus Pseudeucinetus of which Thau- mastodus Champion, 1924 is a junior synonym); an application to the Commission is necessary to maintain usage of Thaumastodinae Champion, 1924 name as valid.” Family euliChadidae Crowson, 1973 Page 297. Delete the entry “*Lichadiden Kolbe, 1908: 249...”. Page 298. Replace “Lichadidae Forbes, 1926: 102” with “Lichadidae Csiki, 1902: 191”. Superfamily elaTeroidea Leach, 1815 Page 298. In the entry “Elaterides Leach, 1815: 85…” replace the “Comment” sec- tion with “Comment: Elateroidea Leach, 1815 given precedence for superfam- ily name over Cebrionoidea Latreille, 1802 (Art. 35.5; ICZN secretariat pers. comm.).” Note. See Kundrata et al. (2019: 87). Family braChypseCTridae Horn, 1881 Page 299. Replace the valid name “Family braChypseCTridae LeConte and Horn, 1883” with “Family braChypseCTridae Horn, 1881”. Page 299. Replace “Brachypsectrini J. L. LeConte and G. H. Horn, 1883: 170” with “Brachypsectrini G. H. Horn, 1881b: 87”. Family euCnemidae Eschscholtz, 1829 Page 301. Replace the entry “Arhipini Cobos, 1965: 396…” with: “Arhipini Cobos, 1965: 396 [stem: Arrhipid-]. Type genus: Arrhipis Bonvouloir, 1871 [as Arhipis, unjustified emendation of type genus name by Fleutiaux (1921: 173), not in prevailing usage; preoccupied genus name, not Arrhipis Agassiz, 1846, un- justified emendation of Arripis Jenyns, 1840 in Pisces]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem forma- tion, not in prevailing usage.” Note. Since Arrhipis Bonvouloir, 1871 is a junior homonym, Reversal of Precedence of the generic name (ICZN 1999: Article 23.9) or an application to the Commission is necessary to conserve usage of this family- group name. Alternatively, a replacement name will be needed. Page 302. At the end of the entry “Epiphanini Muona, 1993: 45…” add “Comment: the senior homonym Epiphanidae Harring, 1913 (type genus Epiphanes Ehren- berg, 1832) in Rotifera is currently used as valid (e.g., Wilts et al. 2012: 180); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Additions and corrections: Family-group names in Coleoptera 87 Page 302. In the entry “Hylocharites Jacquelin du Val, 1859: 119...” replace in the “Com- ment” section “original vernacular name available (Art. 11.7.2): generally accepted as in Muona (2007: 84, as Hylocharini)” with “original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Muona (1993: 43)”. Page 302. In the entries “*Neocharini Muona, 1991a: 167…” and “Neocharini Muona, 1993: 44…” replace “Type genus: Neocharis Sharp, 1887” with “Type genus: Neocharis Sharp, 1877”. Page 303. In the entry “*Galbites Blanchard, 1845b: 71…” replace “Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available in the future then permanently invalid (Art. 39): based on preoccupied type genus.” with “Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized and attributed to Blanchard (1845).” Page 303. Under the entry “*Galbites Blanchard, 1845b: 71…” add: “Gabini [sic] Beaulieu, 1919: 191 [stem: Galb-]. Type genus: Galba Latreille, 1829 [preoccupied genus name, not Galba Schrank, 1803 [Mollusca]; syn. of Galbites Fleutiaux, 1918]. Comment: incorrect original stem formation; permanently inva- lid (Art. 39): based on preoccupied type genus.” Page 305. Above the entry “Macraulacinae Fleutiaux, 1923: 304...” under the valid name “Tribe maCraulaCini Fleutiaux, 1923” add the following entries: “Fornacini Beaulieu, 1919: 191 [stem: Fornac-]. Type genus: Fornax Laporte, 1835. Dromaeolini Beaulieu, 1919: 191 [stem: Dromaeol-]. Type genus: Dromaeolus Kiesenwetter, 1858.” Page 305. At the end of the entry “Macraulacinae Fleutiaux, 1923: 304…” under the valid name “Tribe maCraulaCini Fleutiaux, 1923” add “Comment: Forna- cini Beaulieu, 1919 and Dromaeolini Beaulieu, 1919 take precedence over the valid subfamily and tribe names Macraulacinae/-ini Fleutiaux, 1923, an appli- cation to the Commission is necessary to preserve usage of Macraulacinae/-ini Fleutiaux, 1923 as valid.” Page 305. In the entry “Fornaxini Cobos, 1965: 294...” add at the end of the “Com- ment” section “; family-group name proposed as new without reference to Forna- cini Beaulieu, 1919”. Page 305. Delete the entry “Dromaeolini Leiler, 1976: 48…” Page 306. In the entry “Nematodini Leiler, 1976: 48…” replace “Type genus: Nem- atodes Berthold, 1827” with “Type genus: Nematodes Guérin-Méneville, 1827.” Note. The name Nematodes is usually credited to Berthold (1827: 335). However, it was made available first by Guérin-Méneville in Volume 11 (p. 498) of Bory de Saint-Vincent’s Dictionnaire classique d’histoire naturelle published on January 1827 (title page). The earliest known date of publication for Berthold’s publication is 22 September 1827 (Bousquet 2016: 73). Family ThrosCidae laporTe, 1840 Page 306. In the entry “Stereolia Rafinesque, 1815: 112...” replace “[unjustified emendation of Throscus Latreille, 1797 not in prevailing usage;” with “[unneces- sary replacement name for Throscus Latreille, 1797;”. Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)88 Family elaTeridae Leach, 1815 Pages 306–307. In the entry “Elaterides Leach, 1815: 85…” replace the “Com- ment” section with “Comment: Elateridae Leach, 1815 given precedence for family name over Cebrionidae Latreille, 1802 (Art. 35.5; ICZN secretariat pers. comm.).” Note. See Kundrata et al. (2019: 87). Page 309. In the entry “Oophoridae Gistel, 1848: [5]…” replace “Type genus: Oo- phorus Eschscholtz, 1833” with “Type genus: Oophorus Dejean, 1833.” Page 310. In the entry “Nyctophyxina C. Costa, 1975: 85…” replace “Type genus: Nyctophysis C. Costa, 1975” with “Type genus: Nyctophyxis C. Costa, 1975”. Page 312. In the entry “Campyloxeninae C. Costa, 1975: 114…” replace “Type ge- nus: Campyloxenus Fairmaire, 1860” with “Type genus: Campyloxenus Fairmaire and Germain, 1860”. Page 313. At the end of the entry “Limoniina Jakobson, 1913: 755…” add “Com- ment: the senior homonym Limoniidae/-inae Speiser, 1909 (type genus Limonia Meigen, 1803) in Diptera is currently used as valid (e.g., Ribeiro and Amorin 2002: 1); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 316. Replace the valid name “Subtribe loebliquasimusina Schimmel and Tarnawski, 2009” with “Subtribe loebliquasina Schimmel and Tarnawski, 2009” Page 316. In the entry “Loebliquasina Schimmel and Tarnawski, 2009: 18...” delete the “Comment” section. Page 316. Replace the valid name “Subtribe sTriaToquasimusina Schimmel and Tarnawski, 2009” with “Subtribe sTriaToquasimina Schimmel and Tarnawski, 2009”. Page 316. In the entry “Striatoquasina Schimmel and Tarnawski, 2009: 20...” re- place the stem with “Striatoquasim-” Page 316. Replace the valid name “Subtribe WiTTmeroquasimusina Schimmel and Tarnawski, 2009” with “Subtribe WiTTmeroquasimina Schimmel and Tarnaw- ski, 2009”. Page 316. In the entry “Wittmeroquasina Schimmel and Tarnawski, 2009: 20...” replace the stem with “Wittmeroquasim-” and the type-genus with “Wittmeroquasi- mus Dolin, 1993”. Page 319. Replace the entry “Dicronychidae Schwarz, 1897: 11…” with: “Dicronychidae Schwarz, 1897: 11 [stem: Dicronych-]. Type genus: Dicronychus sensu Laporte, 1840 [not Dicronychus Brullé, 1832; syn. of Eudicronychus Méquig- non, 1931]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).” Page 319. Replace the valid name “Subfamily morosTomaTinae Dolin, 2000” with “Subfamily morosTominae Dolin, 2000”. Page 319. In the entry “Morostominae Dolin, 2000: 18...” replace the stem with “Morostom-” and replace the “Comment” section with “Comment: incorrect origi- nal stem formation maintained under Art. 29.4 (should be Morostomat-).” Additions and corrections: Family-group names in Coleoptera 89 Family lyCidae laporTe, 1836 Page 321. In the entry “Libnetinina Bocák and Bocáková, 1990: 652…” replace “Type genus: Libnetis C. O. Waterhouse, 1878.” with “Type genus: Libnetis C. O. Waterhouse, 1878 [incorrect subsequent spelling of Libnetus by C. O. Waterhouse (1879: 77), incorrect subsequent spelling in prevailing usage, treated as correct original spelling (Art. 33.3.1)].” Page 321 Replace the entry “Lycoprogenthini Bocák and Bocáková, 2008: 709…” with:“Lycoprogenthini Bocák and Bocáková, 2008: 709 [stem: Lycoprogenth-]. Type genus: Lycoprogenthes Pic, 1915 [incorrect subsequent spelling of Lycopro- gentes Pic (1915b: 6), incorrect subsequent spelling in prevailing usage, treated as correct original spelling (Art. 33.3.1)].” Page 322. In the entry “Paralycinae L. N. Medvedev and Kazantsev, 1992: 59…” replace “Type genus: Paralycus L. N. Medvedev and Kazantsev, 1992 [syn. of Ly- ropaeus C. O. Waterhouse, 1878].” with “Type genus: Paralycus L. N. Medvedev and Kazantsev, 1992 [preoccupied genus name, not Paralycus Womersley, 1944 [Acarina]; syn. of Lyropaeus C. O. Waterhouse, 1878]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.” Note. The name Paralycus Womersley, 1944 is not listed in Neave’s Nomenclator Zoologicus. It was made avail- able in Transactions of the Royal Society of South Australia 68: 135. Family lampyridae Rafinesque, 1815 Page 327. Replace the valid name “Tribe CraTomorphini Greene, 1948” with “Tribe CraTomorphini Olivier, 1911”. Page 327. Replace “Cratomorphi Green, 1948: 68, in key” with “Cratomorphinae Olivier, 1911: 75”. Page 327. In the entry “Dadophorini E. Olivier, 1907: 26…” replace “Type genus: Dadophora E. Olivier, 1907” with “Type genus: Dadophora Duponchel, 1844.” Note. The genus Dadophora is credited to Olivier (1907: 27) in the literature but the generic name was first made available by Duponchel (1844: 574) when he described the species Dadophora hyalina. Page 328. Replace the entry “Photini J. L. LeConte, 1881: 30…” with:“Photini J. L. LeConte, 1881: 30 [stem: Photin-]. Type genus: Photinus Laporte, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 2018)]. Comment: placed on the Official List of Family-Group Names in Zoology (as Photinini LeConte, 1881) and correct stem of senior homonym (based on the type genus Photina Bur- meister, 1838) in Mantodea determined to be Photina- to remove the homonymy (ICZN 2018).” Page 329. In the entry “Photurides Lacordaire, 1857: 338…” replace “Type genus: Photuris J. L. LeConte, 1851” with “Type genus: Photuris Dejean 1833”. Family CanTharidae Imhoff, 1856 (1815) Page 331. At the end of the entry “Ichthyurini Champion, 1915: 128…” add the following: “Comment: the senior homonym Ichthyurinae Packard, 1895 (type genus Ichthyura Hübner, 1819) in Lepidoptera is currently a junior synonym of Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)90 Pygaerinae Duponchel, 1845 (see Schintlmeister 2013: 17); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Series ELATERIFORMIA Page 331. Above the valid name “Subfamily CydisTinae Paulus, 1972” add the fol- lowing: “Elateriformia incertae sedis”. Note. The unassigned family-group taxa Cydistinae Paulus, 1972, Pterotinae LeConte, 1861, Ototretinae McDer- mott, 1964, Ototretadrilinae Crowson, 1972, and †Lasiosynidae Kirejtshuk, Chang, Ren and Kun, 2010 belong to Elateriformia incertae sedis (Lawrence et al. 2010a) and are not actually considered subfamilies of Cantharidae. Family JaCobsoniidae Heller, 1926 Page 333. In the entry “Derolathriinae Sen Gupta, 1979: 692…” replace “Type genus: Derolathrus Sharp, 1900” with “Type genus: Derolathrus Sharp, 1908”. Family bosTriChidae Latreille, 1802 Page 335. Replace the valid name “Subfamily dysidinae Lesne, 1921” with “Sub- family dysidinae Lesne, 1894” Page 335. Replace “Dysididae Lesne, 1921b: 286” with “Dysidini Lesne, 1894: 20” Page 337. Under the entry “Lyctides Billberg, 1820a: 48…” add: “Xylotrogidae Schönfeldt, 1887: 128 [stem: Xylotrog-]. Type genus: Xylotrogus Ste- phens, 1830.” Page 337. Move the entry “Tristariini Lesne, 1921b: 287…” above the entry “Tro- goxylini Lesne, 1921a: 231…” and, at the end of the entry “Tristariini Lesne, 1921b: 287…”, add “Comment: Tristariini was proposed before Trogoxylini since Lesne (1921a: 231) included the proper citation and page of Tristariini; in fact, Lesne unnecessarily changed his name Tristariini to Trogoxylini simply because Trogoxylon LeConte, 1862 was older than Tristaria Reitter, 1878; however, we recommend that an application be submitted to the Commission to conserve usage of the well-established name Trogoxylini Lesne, 1921.” Family pTinidae Latreille, 1802 Page 338. Above the valid name “Subfamily pTininae Latreille, 1802” add: “†Subfamily mesernobiinae Engel, 2010 Mesernobiinae Engel, 2010: 32 [stem: Mesernobi-]. Type genus: Mesernobius Engel, 2010.” Page 338. In the entry “Meziini Bellés, 1985: 37, in key…” replace “Type genus: Mezium Curtis, 1828” with “Type genus: Mezium Samouelle, 1819.” Note. The generic name Mezium is credited to Curtis in 1828 in the literature but the name was first made available by Samouelle in 1819 (p. 180) with Ptinus sulcatus Fab- ricius, 1781 as type species by original designation. Page 338. Replace “Gnostidae Gemminger and Harold, 1868: 700” with “Gnosti- dae King, 1866: 317”. Additions and corrections: Family-group names in Coleoptera 91 Page 339. Replace the entry “Cosmoceroideos Solier, 1849: 476...” with: “*Cosmoceroideos Solier, 1849: 476 [stem: Cosmocer-]. Type genus: Cosmocerus Solier, 1849 [preoccupied genus name, not Cosmocerus Guérin-Méneville, 1844 [Coleoptera: Cerambycidae]; syn. of Cerocosmus Gemminger, 1873]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently latinized and at- tributed to Solier (e.g., Lawrence and Newton (1995: 864), as Cosmocerinae) but not generally accepted as valid; if found to be available, permanently invalid (Art. 39): based on preoccupied type genus.” Page 339. Replace “Ernobiinae Pic, 1912: 12” with “Ernobiinae Pic, 1912b: 55.” Page 342. In the entry “Fabiinae Martínez and Viana, 1964: 7…” replace “Type ge- nus: Fabia Martínez and Viana, 1964” with “Type genus: Fabia Martínez and Viana, 1964 [preoccupied genus name, not Fabia Dana, 1851 [Crustacea]; syn. of Fabrasia Martínez and Viana, 1965]” and add “Comment: permanently invalid (Art. 39): based on preoccupied type genus.” Page 342. Below the entry “Fabiinae Martínez and Viana, 1964: 7…” add: “Fabrasiinae Lawrence and Reichardt, 1966: 32 [stem: Fabrasi-]. Type genus: Fabrasia Martínez and Viana, 1965. Comment: replacement name for Fabiinae Martínez and Viana, 1964 because of the homonymy of the type genus.” Family lymexylidae Fleming, 1821 Page 342. In the entry “Hylecoeti Germar, 1818: 344…” replace “Type genus: Hyl- ecoetus Latreille, 1806” with “Type genus: Hylecoetus Latreille, 1806 [syn. of Elat- eroides Schaeffer, 1777].” Family TroGossiTidae Latreille, 1802 Page 343. Replace the valid names “Subfamily pelTinae Latreille, 1806” and “Tribe pelTini Latreille, 1806” with “Subfamily pelTinae Kirby, 1837” and “Tribe pel- Tini Kirby, 1837” respectively. Page 343. Below the valid name “Subfamily pelTinae Kirby, 1837” replace the entry “Peltides Latreille, 1806: 8…” with: “Peltidae Kirby, 1837: 104 [stem: Pelt-]. Type genus: Peltis Kugelann, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].” Page 343. At the end of the entry “Decamerinae Crowson, 1964: 287…” add “Com- ment: the junior homonym Decameridae Rasmussen, 1978 (type genus Decam- eros d’Orbigny, 1850) is available in Echinodermata; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 343. In the entry for “Peltides Latreille, 1806: 8…” under the valid name “Tribe pelTini Kirby, 1837” add an asterisk (*) in front of the name and replace the comment with the following: “Comment: family-group name unavailable (Art. 11.7.1.1): not based on a genus used as valid at the time (see Lawrence and New- ton 1995: 868).” Page 343. Below the entry “*Peltides Latreille, 1806: 8…” add: Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)92 “Peltidae Kirby, 1837: 104 [stem: Pelt-]. Type genus: Peltis Kugelann, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].” Page 344. Replace the valid name “Tribe CaliTyini Reitter, 1922” with “Tribe Cali- Tyini Houlbert, 1922”. Page 344. Replace “Calityni Reitter, 1922a: 66” with “Calityni Houlbert, 1922a: 103” and add at the beginning of the “Comment” section “issued by 15 July 1922; this family-group name was also used the same year by Reitter (1922a [“31 De- cember 1922”] 66, as Calityni;”. Page 344. In the entry “Gymnochilides Lacordaire, 1854b: 344…” replace “Type genus: Gymnochila Klug, 1844” with “Type genus: Gymnochila Erichson, 1844 [syn. of Gymnocheilis Dejean, 1835].” Page 345. Replace the valid name “†Tribe liThosTomaTini Kolibáč and Huang, 2008” with “†Tribe liThosTomini Kolibáč and Huang, 2008”. Page 345. In the entry “Lithostomini Kolibáč and Huang, 2008: 142...” replace the stem with “Lithostom-” and replace the “Comment” section with “Comment: incor- rect original stem formation maintained under Art. 29.4 (should be Lithostomat-).” Family ChaeTosomaTidae Crowson, 1952 Page 345. Replace the entry “Chaetosomatidae Crowson, 1952: 66…” with: “Chaetosomatidae Crowson, 1952: 66 [stem: Chaetosomat-]. Type genus: Chae- tosoma Westwood, 1851 [the senior homonym Chaetosoma Chevrolat, 1843 in Cerambycidae was recently suppressed for both the Principle of Priority and the Principle of Homonymy, placed on the Official Index of Rejected and Invalid Generic Names in Zoology, and Chaetosoma Westwood, 1851 placed on the Of- ficial List of Generic Names in Zoology (ICZN 2011b)]. Comment: the senior homonym Chaetosomatidae Claus, 1872 (type genus Chaetosoma Claparède, 1863) in Nematoda was recently suppressed for both the Principle of Priority and the Principle of Homonymy, placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology and Chaetosomatidae Crowson, 1952 placed on the Official List of Family-Group Names in Zoology (ICZN 2011b).” Family Cleridae Latreille, 1802 Page 346. In the entry “Cylidrina Reitter, 1894: 38…” replace “Type genus: Cylidrus Latreille, 1829” with “Type genus: Cylidrus Latreille, 1817.” Note. The genus- group name Cylidrus was first made available by Latreille (1817: 43). Page 347. Replace the entry “*Trichodites Blanchard, 1845b: 84…” with: “Trichodidae Streubel, 1839: 136 [stem: Trichod-]. Type genus: Trichodes Herbst, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: the younger name originally proposed as Trichodina van der Hoeven, 1849 (type genus Trichoda Müller, 1773) is available in Protozoa: Ciliophora; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Page 347. Delete the entry “Trichodini Portevin, 1931: 457, in key…” Page 348. In the entry “Necrobiaeidae Gistel, 1848: [6]...” replace “Type genus: Necrobia A. G. Olivier, 1795 [placed on the Official List of Generic Names in Additions and corrections: Family-group names in Coleoptera 93 Zoology (ICZN 1961c)]” with “Type genus: Necrobia Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1961c) as “Necrobia A. G. Olivier, 1795”].” Note. The genus-group name Necrobia was first made available by Latreille (1797: 35), not by Olivier since Olivier’s section Nécrobie [No 76bis] in the fourth volume of his Entomologie, ou histoire naturelle des insectes was issued in 1800, not in 1795 (Bousquet 2018: 140). Page 348. In the entry “Dermestoidini Jakobson, 1911a: 719…” replace “Type genus: Dermestoides Schaeffer, 1771” with “Type genus: Dermestoides Schaeffer, 1777”. Family melyridae Leach, 1815 Page 350. In the entry “Pelecophorini Majer, 1987: 797, in key…” replace “Type genus: Pelecophora Lepeletier and Audinet-Serville, 1825” with “Type genus: Pelecophora Dejean, 1821.” Page 352. Replace the entry “*Troglopates Mulsant and Rey, 1867b: 281…” with: “Troglopates Mulsant and Rey, 1867b: 281 [stem: Troglop-]. Type genus: Troglops Erichson, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Porta (1929: 84) and generally accepted as in Mayor (2007: 451, as Troglopini).” Page 353. In the entry “Laiina Jakobson, 1911a: 688…” replace “Type genus: Lai- us Guérin-Méneville, 1838” with “Type genus: Laius Guérin-Méneville, 1830.” Note. The generic name Laius Guérin-Méneville was made available through an illustration (pl. 2, fig. 10) of Laius cyaneus issued in November 1830. Family byTuridae Gistel, 1848 Page 354. Below the entry “Byturidae Gistel, 1848: [3]...” under the valid name “by- Turinae Gistel, 1848” add: “Homoeoplastidae Gistel, 1856a: 360 [stem = Homoeoplast-]. Type genus: Homoeo- plastus Gistel, 1856 [Gistel (1856a: 360) included two available species under the generic name Homoeoplastus, tomentosus, which refers to Dermestes tomentosus De- Geer, 1774, and fumatus, which refers to Dermestes fumatus Linnaeus, 1767; we here select Dermestes tomentosus DeGeer, 1774 as type species; syn. nov. of Byturus Latreille, 1797].” Family sphindidae Jacquelin du Val, 1860 Page 355. In the entry “Odontosphindini Sen Gupta and Crowson, 1979: 180, in key…” replace “Type genus: Odontosphindus Sen Gupta and Crowson, 1979” with “Type genus: Odontosphindus J. L. LeConte, 1878.” Family eroTylidae Latreille, 1802 Page 358. In the entry “Encaustini Crotch, 1876: 476...” replace in the “Comment” section “published after February 1876” with “published in February 1876” Page 358. Replace the entry “Triplacinae Erichson, 1847a: 179…” with: “Triplacina Streubel, 1839: 135 [stem: Triplac-]. Type genus: Triplax Herbst, 1793.” Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)94 Family CrypTophaGidae Kirby, 1826 Page 360. Replace “Paramecosomina Reitter, 1875: 4” with “Paramecosomini Reit- ter, 1874: 381”. Page 360. Replace the valid name “Tribe piCroTini Crowson, 1980” with “Tribe piCroTini Sen Gupta and Crowson, 1971”. Page 360. Replace “Picrotini Crowson, 1980: 283” with “Picrotinae Sen Gupta and Crowson, 1971: 30”. Family phalaCridae Leach, 1815 Page 364. In the entry “Eustilbini Guillebeau, 1892: 149…” replace “Type genus: Eustilbus Sharp, 1888” with “Type genus: Eustilbus Sharp, 1888 [syn. of Stilbus Seidlitz, 1872].” Family laemophloeidae Ganglbauer, 1899 Page 364. Move the entry “Nartheciinae Grouvelle, 1908: 453…” above the entry “Laemophloeini Ganglbauer, 1899: 606…” and replace it with: “Nartheciini Casey, 1890: 497 [stem: Nartheci-]. Type genus: Narthecius J. L. Le- Conte, 1861. Comment: this name is older than the well-established family name Laemophloeidae Ganglbauer, 1899; an application to the Commission is neces- sary to preserve usage of Laemophloeidae Ganglbauer, 1899 over Nartheciidae Casey, 1890.” Family niTidulidae Latreille, 1802 Page 367. Replace the valid name “arhinini Kirejtshuk, 1987” with “arhinopini Kirejtshuk and Bouchard, 2018”. Page 367. In the entry “Arhinini Kirejtshuk, 1987: 63…” replace “Type genus: Arhi- na Murray, 1876” with “Type genus: Arhina Murray, 1867 [preoccupied genus name, not Arhina Agassiz, 1846 (an unjustified emendation for Arina Robineau- Desvoidy, 1830) [Diptera]; syn. of Arhinops Kirejtshuk and Bouchard, 2018]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.” Page 367. Below the entry “Arhinini Kirejtshuk, 1987: 63…” add: “Arhinopini Kirejtshuk and Bouchard, 2018: 157 [stem: Arhinop-]. Type genus: Arhi- nops Kirejtshuk and Bouchard, 2018.” Page 367. Replace “Pityophagini Reitter, 1891: 163” with “Pityophagini Schilsky, 1888: 60”. Page 368. Replace “Glischrochilini Iablokoff -Khnzorian, 1966: 314” with “Glis- chrochilini Chagnon, 1934: 309, in key”. Family Cerylonidae Billberg, 1820 Page 369. Below the entry “Tachyoryctidiini Jeannel and Paulian, 1945…” add: “Hypodacninae Dajoz, 1976: 184, in key [stem: Hypodacn-]. Type genus: Hypodacne J. L. LeConte, 1875.” Page 370. Above the entry “Murmidiides Jacquelin du Val, 1858: 227…” add: Additions and corrections: Family-group names in Coleoptera 95 “Ceuthocera Mannerheim, 1852: 383 [stem: Ceutocer-]. Type genus: Ceutocerus Germar, 1823 [as Ceuthocerus, incorrect subsequent spelling of type genus name, not in prevailing usage; syn. of Murmidius Leach, 1822]. Comment: incorrect original stem formation, not in prevailing usage; the discovery of this family- group name threatens the well-established name Murmidiinae Jacquelin du Val, 1858; as far as we know, a family-group name based on Ceutocerus has not been used as valid after 1899 and we found 25 references, published by at least 10 authors in the immediately preceding 50 years and encompassing a span of not less than 10 years, using the family-group name based on Murmidius as valid; therefore Reversal of Precedence (Art. 23.9) is used (see Appendix 1 for support- ing references) to qualify Murmidiinae as nomen protectum and Ceutocerinae as nomen oblitum. Family endomyChidae Leach, 1815 Page 372. Below the valid name “Subfamily anamorphinae Strohecker, 1953” add: “Symbiotinae Joy, 1932: 558, in key [stem: Symbiot-]. Type genus: Symbiotes Redten- bacher, 1849.” Page 372. At the end of the entry “Anamorphini Strohecker, 1953: 15, in key…” add “Comment: the genus Symbiotes Redtenbacher, 1849 is currently placed in the subfamily Anamorphinae Strohecker, 1953 (e.g., Tomaszewska 2007: 559), therefore the older name Symbiotinae Joy, 1932 takes precedence over Anamo- rphinae based on the Principle of Priority; an application to the Commission is necessary to preserve usage of Anamorphinae Strohecker, 1953 as valid.” Page 373. In the entry “Epipocidae Gorham, 1873: 20…” replace “Type genus: Epipo- cus Germar, 1843” with “Type genus: Epipocus Chevrolat, 1836”. Page 373. In the entry “Corynomalidae Gorham, 1873: 14…” replace “Type genus: Corynomalus Gerstaecker, 1857” with “Type genus: Corynomalus Chevrolat, 1836”. Family CoCCinellidae Latreille, 1807 Page 374. Replace the valid name “Tribe seranGiini Pope, 1962” with “Tribe seranGiini Blackwelder, 1845”. Page 374. In the entry “*Serangiini Blackwelder, 1945: 450…” remove the asterisk (*) and replace the “Comment” section with “Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Pope (1962: 627, as Serangiini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (Art. 13.2.1).” Page 374. Delete the entry “Serangiini Pope, 1962: 627…” Page 376. In the entry “*Mysiates Mulsant, 1846: 125…” replace “[stem: Myzi-]. Type genus: Myzia Mulsant, 1846 [as Mysia, alternative original spelling of type genus name; we follow Kovář (2007: 620) in using Myzia as the correct spelling for this genus]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.” Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)96 with “[stem: Mysi-]. Type genus: Mysia Mulsant, 1846. Comment: original vernac- ular name unavailable (Art. 11.7.2): not subsequently latinized.” Note. Mulsant (1846) used two original spellings, Myzia and Mysia. Subsequently Mulsant (1850: 137) used the spelling Mysia and so acted as First Reviser (Art. 24.2.4); Mysia is therefore considered the correct original spelling. Page 377. Below the entry “Synonychini Weise, 1885: 7…” add: “Hippodamiini Weise, 1885: 11 [stem: Hippodami-]. Type genus: Hippodamia Chev- rolat, 1836.” Page 377. Above the entry “Anisolemniina Mader, 1954: 93, in key…” add: “Cheilomenini F. A. Schilder and M. Schilder, 1928: 218 [stem: Cheilomen-]. Type genus: Cheilomenes Chevrolat, 1836. Veraniini F. A. Schilder and M. Schilder, 1928: 218 [stem: Verani-]. Type genus: Verania Mulsant, 1850 [preoccupied genus name, not Verania Krohn, 1846 [Mol- lusca]; syn. of Micraspis Chevrolat, 1836]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.” Page 378. In the entry “Subcoccinellini Jakobson, 1915: 968…” replace “Type ge- nus: Subcoccinella Huber, 1841” with “Type genus: Subcoccinella Agassiz, 1846.” Note. Huber (1841) proposed the new genus-group name in a vernacular form only (i.e., Subcoccinelle) and therefore the name is unavailable. Agassiz (1846a: 156) is the first who used the name as valid, latinized it, and provided a reference to a previously published description. Page 379. At the end of the entry “Monocorynini Miyatake, 1988: 28…” add: “Comment: the family group name Monocoryninae Rees, 1956 (type genus Monocoryne Broch, 1910) is available in Cnidaria; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).” Family Corylophidae LeConte, 1852 Page 382. In the entry “Periptyctinae Ślipiński et al., 2001: 312…” replace “Type ge- nus: Periptyctus Blackburn, 1825” with “Type genus: Periptyctus Blackburn, 1895.” Page 382. Replace “Clypeastres L. Redtenbacher, 1845: 122” with “Clypeastrina Perty, 1840: 928”. Page 383. Replace the valid name “Tribe seriCoderini Matthews, 1888” with “Tribe seriCoderini Matthews, 1886”. Page 383. Replace “Sericoderina A. Matthews, 1888: 103” with “Sericoderina A. Matthews, 1886: 224”. Family Akalyptoischiidae Lord, Hartley, Lawrence, McHugh, Whiting and Miller, 2010 Page 383. Replace the valid name “Family aKalypToisChiidae Lord, Hartley, Law- rence, McHugh and Miller, 2010” with “Family aKalypToisChiidae Lord, Hart- ley, Lawrence, McHugh, Whiting and Miller, 2010”. Family laTridiidae Erichson, 1842 Page 384. Under the valid name “Family laTridiidae Erichson, 1842” replace the entry “Lathridien Erichson, 1842: 122…” with: Additions and corrections: Family-group names in Coleoptera 97 “Lathridien Erichson, 1842: 122 [stem: Latridi-]. Type genus: Latridius Herbst, 1793 [placed on the Official List of Generic Names in Zoology (ICZN 2011c)]. Com- ment: family name Latridiidae given precedence over Corticariidae Curtis, 1829 and other family-group names based on Corticaria Marsham, 1802 when- ever their type genera are placed in the same family-group taxon, and placed on the Official List of Family-Group Names in Zoology (ICZN 2011c).” Page 384. Under the valid name “Subfamily laTridiinae Erichson, 1842” replace the entry “Lathridien Erichson, 1842: 122…” with: “Lathridien Erichson, 1842: 122 [stem: Latridi-]. Type genus: Latridius Herbst, 1793 [placed on the Official List of Generic Names in Zoology (ICZN 2011c)]. Com- ment: original vernacular name available (Art. 11.7.2): first used in latinized form by L. Redtenbacher (1845: 123, as Lathridii), generally accepted as in Lawrence and Newton (1995: 886, as Latridiidae); incorrect original stem formation, not in prevailing usage.” Note. The name “Latridii” used earlier by Westerhauser (1832: 151) is considered a plural term for the members of the genus Latridius, Wester- hauser’s name is therefore unavailable as a family-group name (Art. 11.7.1.2). Page 384. Replace the entry “Corticaridae Curtis, 1829: pl. 283…” with: “Corticaridae Curtis, 1829: pl. 283 [stem: Corticari-]. Type genus: Corticaria Mar- sham, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2011c)]. Comment: incorrect original stem formation, not in prevailing usage; Corticaridae Curtis, 1829 placed on the Official List of Family-Group Names in Zoology, with the endorsement that it and other family-group names based on Corticaria are not to be given priority over Latridiidae Erichson, 1842 and other family-group names based on Latridius Herbst, 1793 whenever their type genera are placed in the same family-group taxon (ICZN 2011c).” Page 384. Replace the entry “*Melanophthalmidae Arnett, 1962b: 835…” with: “Melanophthalmidae Stickney, 1923: 45 [stem: Melanophthalm-]. Type genus: Mel- anophthalma Motschulsky, 1866.” Page 384. Replace the valid name “†Subfamily TeTrameropseinae Kirejtshuk and Azar, 2008” with “†Subfamily TeTrameropsinae Kirejtshuk and Azar, 2008” Page 384. In the entry “Tetrameropsinae Kirejtshuk and Azar, 2008: 36...” replace the stem with “Tetramerops-” and replace the “Comment” section with “Comment: incor- rect original stem formation maintained under Art. 29.4 (should be Tetrameropse-).” Family myCeTophaGidae Leach, 1815 Page 385. Replace “Triphyllini Harold, 1880: 757” with “Triphyllidae Crotch, 1873a: 82”. Family Ciidae Leach, 1819 Page 386. Below the entry “Cisidae Leach, 1819: 206…” under the valid name “Tribe Ciini Leach, 1819” add: “Ennearthroninae Chûjô, 1939a: 9 [stem: Ennearthr-]. Type genus: Ennearthron Mel- lié, 1847. Comment: Ennearthroninae was also used later in the same year by Chûjô (1939b: 24, in key); incorrect original stem formation, not in prevailing usage.” Patrice Bouchard & Yves Bousquet / ZooKeys 922: 65–139 (2020)98 Family mordellidae Latreille, 1802 Page 390. Above the valid name “Tribe mordellini Latreille, 1802” add: “Tribe CurTimordini Odnosum, 2010 Curtimordini Odnosum, 2010: 69, 119 [stem: Curtimord-]. Type genus: Curtimorda Méquignon, 1946.” Page 390. Above the valid name “Tribe reynoldsiellini Franciscolo, 1957” add: “Tribe mordelloChroini Odnosum, 2010 Mordellochroini Odnosum, 2010: 71, 125 [stem: Mordellochro-]. Type genus: Mordellochroa Emery, 1876.” Family ripiphoridae Laporte, 1840 Page 390. Replace the valid name “Family ripiphoridae Gemminger, 1870 (1855)” with “Family ripiphoridae Laporte, 1840”. Page 390. Replace the entry “Rhipiphoridae Gemminger, 1870: 2117…” with: “Rhipiphorites Laporte, 1840: 261 [stem: Ripiphor-]. Type genus: Ripiphorus Bosc, 1791 [as Rhipiphorus, unjustified emendation of type genus name by Duméril (1827: 374), not in prevailing usage].” Page 391. Replace the valid name “Subfamily peleCoTominae Seidlitz, 1875” with “Subfamily peleCoTominae Guérin-Méneville, 1857”. Page 391. Replace “Pelecotomini Seidlitz, 1875 [Gatt.]: 104” with “Pelecotomini Guérin-Méneville, 1857: 91”. Page 391. Replace the valid name “Subfamily ripiphorinae Gemminger, 1870 (1855)” with “Subfamily ripiphorinae Laporte, 1840”. Page 391. Replace the entry “Rhipiphoridae Gemminger, 1870: 2117…” with: “Rhipiphorites Laporte, 1840: 261 [stem: Ripiphor-]. Type genus: Ripiphorus Bosc, 1791 [as Rhipiphorus, unjustified emendation of type genus name by Duméril (1827: 374), not in prevailing usage].” Page 391. Below the valid name “Tribe maCrosiaGonini Heyden, 1908” delete the entry “Rhipiphorites Laporte, 1840b: 261…” Page 392. In the entry “Macrosiagonini L. Heyden, 1908: 45…” delete the “Com- ment” section. Page 392. Replace the valid name “Tribe ripiphorini Gemminger, 1870 (1855)” with “Tribe ripiphorini Laporte, 1840”. Page 392. In the entries “*Mioditini A. Costa, 1853: 2…” and “Myoditini Ger- staecker, 1855: 15…” replace “Type genus: Myodites Latreille, 1829” with “Type genus: Myodites Latreille, 1819”. Page 392. In the entry “Myoditini Gerstaecker, 1855: 15…” delete the “Comment” section. Page 392. Replace the entry “Rhipidophoridae Gemminger, 1870: 2117…” with: “Rhipiphorites Laporte, 1840: 261 [stem: Ripiphor-]. Type genus: Ripiphorus Bosc, 1791 [as Rhipiphorus, unjustified emendation of type genus name by Duméril (1827: 374),